Development Summary

 

Cells commit to cell fates, which can be redefined, thus making development flexible

Regulation is integral in development

 

The xx/xy system of sex determination is believed to have arisen independently many times over

 

The germ line is a binary, irreversible decision that happens in early development

P granules and the cell cytoskeleton contribute to the separation of soma from germ line in early development.

Localization of mRNAs within a cell is accomplished by anchoring the mRNAs to one end of polarized cytoskeleton chains.

The formation of concentration gradients of the two transcription factors BCD and HB-M allows positional information along the A-P axis to be established.
Positional information can be established by cell-cell signaling through a concentration gradient of a secreted signaling molecule.


Through transcription-factor regulation patterns, positional information leads to the formation of the correct number of segments.
The complexity of regulatory elements of the pair-rule genes turns an asymmetric (gap-gene) expression into a repeating one.


Segement identity is established through asymmetric gap-gene expression patterns that deploy an asymmetric pattern of homeotic gene expression

 

Intracellular and intercellular positive feedback loops make sure the fate of cells is remembered.
Components of developmental pathways are mixed and matched to make new pathways.

Insect and mammalian homeotic genes are arranged on the chromosome in an order that reflects their spatial expression from anterior to posterior.
The same basic building blocks and regulatory devices are used in most organisms.
Plant and animal cells use the same strategy to control gene expression and transcription.