Fact: current life forms differ
from those of previous times,
and are 'descended' from them.
Evolution is descent with modification.
["Descent" in the genealogical
sense]
Fact: current life forms are
extremely variable,
both within and among species.
Evolution accounts for variation among living
organisms.
[Variation occurs in time
and space]
"Theory of Evolution"
The explanation
of observed patterns of temporal & spatial variation
in terms of biological and physical processes.
Natural
Selection provides a mechanism for Evolution:
Modern evolutionary theory seeks to clarify this mechanism.
The observable
order in Nature is due to common
descent from an ancestor:
Organisms resemble each other because they are related.
Fact:
All living things are related (the basic fact of biology):
The degree
of relationship provides a basis for "natural
classification":
Taxonomy should reflect the phylogeny of organisms.
I summon the supernatural beings
Who first contrived
The transmorgifications
In the stuff of life.
You did it for your own amusement.
Descend again, be pleased to reanimante
This revival of those marvels.
Reveal, now, exactly
How they were performed
From the beginning
Up to this moment.
Ovid: Metamorphoses (trans. Ted Hughes)
The Classical
Tradition: Plato & Aristotle
Theory
of Forms (essences, eidos)
'real' objects are manifestations of 'ideal' forms
variation is illusory [see Plato "The
Republic"]
Dichotomy:
the world is composed of paired opposites
"A" versus "not A" classes
good / bad, right / wrong, up / down, light / dark, male / female, etc.
e.g., vertebrates vs. invertebrates
Aristotle
(384-322 BCE) - "Father of Biology"
Five books on zoology ("Generation
of Animals")
Biological structures have purpose: Efficient versus Final Causes
Natural
Theology (John Ray
1627-1705)
"The Wisdom of God, Manifested in His Creation"
(1691)
'Ideal' forms exist in the Mind of God:
'real' world created by God (Genesis
1:1)
The study of nature is a pious activity
Scala Naturae: the "Great
Chain of Being"
Creation is an infinitely graduated progressive series
Time scale is short (ca. 6,000 years)
Species are static: no new forms, no change, no extinction
Linnean
Taxonomy (Carl von Linne [Carolus
Linneaus]
1707-1778)
"Systema Naturae" (1735; 10th
ed. 01 January 1758)
4,162 animals described
binomial nomenclature: genus
+ species names
Worked "Ad majorem Dei gloriam": for the greater glory of God
Exploration
creates a Scientific Crisis
New
forms are discovered that don't fit the Scala
Extinctions
have evidently occurred
Variation
is real in space: what about over time?
The Darwinian Revolution
Charles
Darwin
(1809-1882)
B.Sc. (Cambridge): pre-med
Naturalist on board HMS "Beagle" (1831-36)
"The Voyage of the Beagle" (1839) a
best-seller
Read Robert Malthus "On
Population" (1838):
population increases exponentially, resources increase arithmetically
[Married Emma on 29 Jan 1839]
Letter from Alfred Wallace
(1823-1913) in June 1858
Observation: In any species, more young are born than can possibly survive.
Observation: Yet a species' numbers do not increase without limit.
CONCLUSION: There is a Struggle
for Survival,
and differential survival and reproduction occur
within species.
[Darwin: "I use 'struggle' in a large
and metaphorical sense..."].
Observation: Individuals within
species show variation
that affects the probability that they will survive this struggle and
leave offspring.
CONCLUSION: Those individuals that
survive and reproduce do so in consequence
of their
"adaptively superior" variation (they
are "more fit")
This process
of differential survival and reproduction is called Natural
Selection.
Observation: Variation is heritabile:
offspring tend to resemble their parents.
[Remember that Mendelian
genetics was unknown in 1859).
CONCLUSION:
Adaptively superior variation will be inherited by the offspring generation.
That is,
evolution occurs as descent with modification.
For further reading:
Loren Eisley (1959).
"Darwin's Century." Doubleday.
William Irvine (1955).
"Apes, Angels, and Victorians: Darwin, Huxley, & Evolution." McGraw-Hill.
Ernst Mayr (1994).
"One Long Argument". Harvard University Press.
Gordon Ratray Taylor
(1963). "The Science of Life." McGraw-Hill.