Pteridophytes of the Canadian Arctic Archipelago

S.G. Aiken, M.C. LeBlanc, and M.J. Dallwitz


Character List

#1. <Common name>/

#2. <Family>/
1. Asteraceae (Compositae), daisy family/
2. Betulaceae, birch family/
3. Boraginaceae, lungwort family/
4. Brassicaceae (Cruciferae), Draba family/
5. Campanulaceae, bluebell family/
6. Caryophyllaceae, pink family/
7. Crassulaceae, stonecrop family/
8. Cyperaceae, sedge family/
9. Diapensiaceae/
10. Empetraceae, crowberry family/
11. Ericaceae, bilberry family/
12. Equisetaceae, horsetail family/
13. Fabaceae (Leguminosae), pea family/
14. Gentianaceae/
15. Haloragaceae, watermilfoil family/
16. Hippuridaceae, mares tail family/
17. Juncaceae, rush family/
18. Lentibulariaceae, bladderwort family/
19. Liliaceae (Nartheciaceae), Tofieldia family/
20. Linaceae, flax family/
21. Lycopodiaceae, club-moss family/
22. Onagraceae, fireweed family/
23. Orchidaceae, orchid family/
24. Papaveraceae, poppy family/
25. Plantaginaceae, plantain family/
26. Plumbaginaceae, leadwort family/
27. Poaceae, grass family/
28. Polemoniaceae/
29. Polygonaceae, buckwheat family/
30. Polypodiaceae, fern family/
31. Portulaceae/
32. Potamogetonaceae, pondweed family/
33. Primulaceae/
34. Pyrolaceae/
35. Ranunculaceae, buttercup family/
36. Rosaceae, rose family/
37. Salicaceae, willow family/
38. Saxifragaceae, saxifrage family/
39. Scrophulariaceae, fernweed family/
40. Sparganiaceae/

#3. <Place of valid publication>/

#4. Nomenclatural section used by Flora of North America project <relevant to the Cyperaceae>/

Information supplied by Dr. P. Ball, University of Toronto.

#5. <Synonymy based on type specimen>/

#6. Type: <specimen>/

#7. <Synonyms based on different type specimens>/

#8. <Synonymy, according to Kartesz (1994), Hultén, (1968), Porsild (1957), Polunin (1940)>/

Vegetative Morphology

#9. Plants <habit: whether woody>/
1. herbs /
2. shrubs/

HABIT: general appearance, characteristic form, or mode of growth of the plant.

HERB: a plant with no persistent or woody parts above ground.

SHRUB: a perennial woody plant usually with several main stems arising from or near the ground.

#10. Plants <longevity>/
1. annual herbs/
2. biennial herbs/
3. perennial herbs/

HERB: a plant with no woody parts above ground.

ANNUAL: a plant that completes its life cycle, from seed germination to seed production followed by death, within a single growing season.

PERENNIAL: a plant that continues growth from year to year.

#11. Plants <shrub habit>/
1. dwarf shrubs <0.1 m or less, e.g. S. reticulata>/
2. low shrubs <0.15–0.5 m, e.g. S. calcicola>/
3. mid shrubs <0.6–2.0 m, e.g. some S. alaxensis or S. richardsonii>/
4. tall shrubs <greater than 2.0 m, e.g. some S. planifolia or S. pulchra>/
5. trees <sometimes with several trunks, but of "tree" stature, e.g. some S. alaxensis>/

HABIT: general appearance, characteristic form, or mode of growth of the plant.

DWARF SHRUBS are diminutive (very small) plants usually with erect stems. The leaves and catkins may also be diminutive, but not always. Species such as Salix arctica, which is a dwarf shrub, may have dwarf stature or be prostrate but have leaves and catkins that are as large as those of tall shrubs.

#12. Plants <whether caespitose>/
1. caespitose/
2. not caespitose /

CAESPITOSE: with several stems growing tightly together and upward from the base in tufts; tufted.

#13. Plants <general appearance>/

PLANT APPEARANCE: as seen in living plants growing in a typical Arctic habitat and not stressed by pollution or other abnormal envirnomental factor. Be aware that plant appearance is often changed in herbarium specimens. SINGLE-STEMMED: as in an annual plant, or perennial that may only produce one stem per growing season in the Arctic.

LOOSE CLUSTERS: two or more stems growing loosely together and upward from the base. These are often plants with short rhizomes.

TUFTED: or CESPITOSE: with several stems, growing tightly together and upward from the base, in tufts.

CUSHION-LIKE: rounded, compact, plants; stems not visible.

MATTED: flattened, spread out, stems tangled or interwoven.

BASAL ROSETTES: dense radiating clusters of leaves at or near ground level.

CAUDEX: a short main stem zone above a tap root. Plants with an abrupt transition between the root, usually a taproot and the leaves.

BRANCHING STEMS: main stem with distinct branches at intervals along the stem.

TRAILING stems: stems arising from a root zone and growing along the surface of the ground for considerable lengths relative to the size of the leaves.

#14. Plants <size>/
1. less than 15 cm high/
2. more than 15 cm high/

#15. Plants <height>/
cm high/

#16. Plants <vegetative reproduction>/
1. not colonial /
2. forming colonies by layering <e.g. some Salix arctic or Salix arctophila>/
3. forming rhizomatous or stoloniferous mats <e.g. Salix herbacea>/

COLONIAL: groups of plants connected to one another by underground organs.

LAYERING: branches lying on the top of the ground and rooting.

RHIZOMATOUS MATS: mats formed from rhizomes which are horizontal underground stems.

#17. Plants <vegetatively proliferating by bulbils>/
1. vegetatively proliferating by bulbils not in the inflorescence/
2. not vegetatively proliferating by bulbils /

VEGETATIVELY proliferating: adventitious leafy or bulb-like structures developing on the parent plant, often associated with flowering stem or inflorescence.

#18. Plants <whether with milky juice>/
1. with milky juice/
2. without milky juice /

MILKY JUICE: white latex substance.

#19. Plants <whether glandular viscid>/
1. glandular viscid/
2. not glandular viscid /

GLANDULAR VISCID: sticky to the touch when fresh from fluid secreted by glands. GLANDS: small bulges that store or secrete fluids.

Roots

#20. Taproot <presence>/
1. present/
2. absent <roots fibrous> /

TAPROOT: a large primary root with much smaller lateral roots.

#21. <Root comment>/

#22. Roots <external colour, scored dried, young material>/
1. colourless/
2. white/
3. yellow <not yellow brown>/
4. pallid-brown <includes yellow-brown>/
5. red-brown <includes dark-brown and chestnut-coloured>/
6. purple/
7. grey/
8. black/

#23. Caudex <whether present>/
1. present/
2. absent/

CAUDEX: a short main stem zone above a tap root. Plants with an abrupt transition between the root, usually a taproot, and the leaves.

Stems near ground level

#24. Ground-level or under-ground stems <horizontal stems either rhizomes or stolons or with vertical under-ground stem or caudex; character applies to herbs, not shrubs>/
1. horizontal/
2. vertical and often branched <sometimes forming a stout caudex>/
3. not developed horizontally or vertically underground/

HORIZONTAL STEMS: includes rhizomes and stolons. In the Arctic this general term is appropriate as whether horizontal stems are above or below ground level varies with the substrate that plants are growing on. Tufted plants like some grassess and sedges or compact cushion plants have not easily seen horizontal stems.

VERTICAL UNDERGROUND OR GROUND LEVEL STEMS: conspicuous stems development in these positions, usually plants with little above ground stem development. CAUDEX: The persisting transition stem zone between the roots and the otehrwise annual leaves.

Annual plants usually have no horizontal stems.

#25. Ground-level or under-ground stems <horizontal stems whether rhizomes or stolons>/
1. rhizomatous <below ground-level>/
2. stoloniferous <above ground-level>/

RHIZOME: an underground, horizontal stem, usually with reduced scaly leaves and adventitious roots at regularly spaced nodes.

STOLON: a horizontal stem which creeps over the surface of the ground. The end of the stolons root and vegetatively propagate new plants. eg. Spider Saxifrage and Puccinellia ×phryganodes .

#26. Ground-level or under-ground stems <whether elongate>/
1. elongate/
2. compact/

ELONGATE: rhizomes or stolons usually more than 5 cm long.

COMPACT: rhizomes or stolons 1–2 cm long.

#27. Ground-level or under-ground stems <width>/
mm wide/

#28. Scales <present or absent at the base of stems or on rhizomes or stolons>/
1. present/
2. absent/

STEM SCALES: reduced leaf-like structures that usually have a single prominent ridge, unlike prophylls that have two such ridges.

#29. <Number of scale-like leaves> scales/

#30. Scales <on rhizomes or stolons, surface>/
1. smooth/
2. striate/
3. grooved/

RHIZOME SCALES: reduced scale-like leaves on underground, horizontal stems.

SMOOTH: surface devoid of markings or ornamentation (hair types are not considered here).

STRIATE: with fine longitudinal lines or ridges.

GROOVED: with deep longitudinal lines or ridges.

#31. Scales <on rhizomes or stolons, length>/
mm long/

RHIZOME SCALES: reduced scaly leaves on underground, horizontal stems.

#32. Scales <on rhizomes or stolons, whether hairy>/
1. glabrous/
2. hairy/

RHIZOME SCALES: reduced scaly leaves on underground, horizontal stems.

GLABROUS: without hairs.

Stems

#33. Aerial stems <position>/
1. erect <aerial stems>/
2. ascending/
3. decumbent/
4. prostrate <trailing on the ground>/

ERECT: a stem that is vertical from the roots not declining or trailing.

DECUMBENT: a stem that lies flat on the ground but with the tip ascending.

PROSTRATE: a stem that lies flat on the ground.

NOTE: There are many plants in the Arctic that have a basal rosette or tuft of leaves from which a long mostly leafless stem emerges with a terminal flower or inflorescence. This is called a scape or flowering stem to distinguish it from a vegetative (leaf-bearing) stem; the vegetative stems may be very short and should be looked for within the basal rosette or tuft of leaves. In Juncus, an erect stem occurs between the rhizome and the leaf subtending the inflorescence that may be terminal or lateral. Several Arctic species of grasses have been observed to develop flowering stems that are initially prostrate but that become almost erect at anthesis, for example, some species of Puccinellia. These have been scored as having both states.

TRAILING stem lies flat on the ground (prostrate) and creeps but does not root. Term used for willows.

#34. Aerial stems <whether jointed>/
1. conspicuously jointed with nodes covered by whorls of tiny leaves fused for part of their length into sheaths that are tipped with teeth /
2. not conspicuously jointed /

JOINTED STEMS: stems easily pulled apart at the nodes because of a change in the position of the stem vasculature at each node.

#35. Aerial stems <whether filiform> /
1. filiform/
2. not filiform /

FILIFORM: thread-like.

#36. Aerial stems <shape in cross-section, mid-third of the uppermost internode, cf. Metcalfe 1971>/
1. triangular in cross-section/
2. circular or oval in cross-section/
3. squarish/

TRIANGULAR: usually an equilateral triangle, sometimes with rounded corners./par Character mainly used for Cyperaceae.

#37. Aerial stems with <number of>/
ridges /

RIDGES: raised areas that are the result of underlying sclerenchyma separated by zones of less, or no cell sclerenchyma.

Stem vestiture

#38. Aerial stems <hairs>/
1. glabrous/
2. sparsely hairy/
3. densely hairy/
4. scabrous/

In Dicotyledons this character only refers to stems bearing leaves (vegetative stems) which may have flowers borne in leaf axils but not a terminal flower or inflorescence. In grasses this term refers to internodes: the portion of the stem between nodes, often on flowering stems.

GLABROUS: surface smooth without any form of hairs or surface roughness.

HAIRY: with soft hairs that lack stiffness.

SCABROUS: surface with short stiff hairs that make the surface feel rough.

RESINOUS WART-LIKE GLANDS: conspicuous bumpy structures that secret resin. Found in Betula glandulosa.

GLANDULAR HAIRS: hairs that have a bulge that may secrete fluids at the tip.

#39. Aerial stems <whether glandular structures present>/
1. covered with resinous wart-like glands/
2. with glandular hairs/
3. without either glandular hairs or resinous warts /

RESINOUS WART-LIKE GLANDS: Small bulges on the stem that secrete resin.

GLANDULAR HAIRS: hairs that have a bulge that may secrete fluids at the tip.

#40. Aerial stem hairs <orientation>/
1. appressed/
2. spreading/
3. erect/
4. reflexed/

APPRESSED: hairs at angles less than 30ø.

SPREADING: hairs at angles 30–60°.

ERECT: hairs at 90°.

REFLEXED: hairs pointing down at angles more than 90°.

Branches

#41. Branches <colour of 1- to 2-year-old shoots>/
1. yellowish/
2. grey-brown/
3. red-brown/
4. violet/
5. yellow-brown/
6. brownish/

BRANCH is a two year old or older vegetative shoot.

#42. Branches <glaucescence>/
1. not glaucous <lacking waxy coating> /
2. thinly glaucous <visible when scratched or with isolated crystals>/
3. thickly glaucous <bluish or whitish waxy coating>/

BRANCH is a two year old or older vegetative shoot.

GLAUCOUS: covered with a whitish or bluish waxy coating (bloom), as on the surface of purple plums.

#43. Branches <general hairiness>/
1. glabrous/
2. glabrescent/
3. hairy/

BRANCH is a two year old or older vegetative shoot

GLABROUS: without hairs.

GLABRESCENT: initially hairy but becoming sparsely hairy of glabrous(smooth) with age.

HAIRY: with any form of hairs.

#44. Branches <pubescence type>/
1. pilose/
2. villous/
3. tomentose/
4. woolly/
5. with long-silky hairs/

BRANCH is a two year old or older vegetative shoot.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: bearing short, dense, soft, wooly hairs, that are matted or tangled.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

#45. Branches epidermis <flakiness of 3–4-year old shoots>/
1. not flaky /
2. flaky/

BRANCH is a two year old or older vegetative shoot.

EPIDERMIS: the outermost layer of bark; refers to young branches.

FLAKY: peeling in relatively large pieces.

Branchlets

#46. Branchlets <colour>/
1. yellow-green/
2. yellow-brown/
3. grey-brown/
4. red-brown/
5. violet/
6. brownish/

BRANCHLET: the current year's shoot.

#47. Branchlets <glaucescence>/
1. not glaucous <lacking any waxy coating>/
2. thinly glaucous/
3. thickly glaucous/

BRANCHLET: the current year's shoot.

GLAUCOUS: covered with a whitish or bluish waxy coating (bloom), as on the surface of a plum.

THINLY GLAUCOUS: waxy surface visible when scratched or with isolated patches of wax.

THICKLY GLAUCOUS: covered with a bluish or whitish coating,

#48. Branchlets <general hairiness>/
1. glabrous/
2. glabrescent/
3. hairy/

BRANCHLET: the current year's shoot.

GLABROUS: without hairs.

GLABRESCENT: initially hairy but becoming glabrous.

#49. Branchlets <pubescence type>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. woolly/
6. with short-silky hairs <less than 0.5 mm>/
7. with long-silky hairs <greater than 0.5 mm>/

BRANCHLET: the current year's shoot.

GLABROUS: without hairs.

PUBERULENT: minutely pubescent or downy; hairs very short.

PUBESCENT: covered with short, fine hairs, downy.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

#50. Branchlets hairs <density>/
1. sparse/
2. moderately dense <surface 50% visible>/
3. very dense <surface completely obscured>/

BRANCHLET: the current year's shoot.

#51. Branchlets hairs <orientation to surface>/
1. appressed/
2. fishhook-curved/
3. spreading/

BRANCHLET: the current year's shoot.

APPRESSED: hair pressed close or flat against the surface.

FISHHOOK-CURVED: shaped, from the surface, as a fish hook.

SPREADING: hair standing out from the surface.

#52. Bud scale inner membrane <adnation, as seen on open buds>/
1. free and separating from outer membrane/
2. free but not separating from outer membrane/
3. fused to outer layer/

BUD SCALE:

#53. Leaves <presence>/
1. present and green /
2. absent, or reduced and scale-like near the base of the stem/

SCALE-LIKE: resembling a scale, and lacking green colour.

#54. Leaves <description if reduced and scale-like>/
1. merely prophylls <e.g. Juncus arcticus>/
2. merely achlorophyllous sheathing scales <Orchidaceae>/

PROPHYLL: the first leaf of a lateral shoot. It is a leaf sheath with two pronounced veins.

ACHLOROPHYLLOUS: lacking chlorophyll; not green.

Leaves

#55. Leaves <position>/
1. mostly basal/
2. distributed along the stems/
3. arising singly from creeping rhizomes/
4. all basal in a rosette/

MOSTLY BASAL:

vertical leaves clustered at the base of the plant with conspicuously few leaves along the stems.

A BASAL ROSETTE: a dense radiating cluster of leaves at or near ground level.

PROPHYLL:

#56. Leaves <attachment>/
1. alternate/
2. opposite/
3. whorled/

ALTERNATE: a form of leaf arrangement in which there is one leaf at each node.

OPPOSITE: a form of leaf arrangement in which the leaves arise in pairs at each node.

WHORLED: a form of leaf arrangement in which three or more leaves arise at each node.

#57. Leaves <whether distichous>/
1. distinctly distichous/
2. not distinctly distichous /

DISTICHOUS: two ranked; in two rows so that the leaves are in approximately the same plane.

#58. Leaves <simple or compound>/
1. simple /
2. compound <including fern fronds>/

SIMPLE LEAF: a leaf blade that is not separated into leaflets, even though the blade may be deeply lobed or cleft.

COMPOUND LEAF: a leaf blade that is separated into two or more distinct leaflets.

FROND: the leaf-like structure of a fern consisting of a central rachis and pinnae, leaflet-like blades that may be further pinnately divided.

#59. Leaves <heterophyllous>/
1. heterophyllous /
2. not heterophyllous /

HETEROPHYLLOUS: having different shapes or textures of leaves on the same plant.

#60. Leaves <longevity>/
1. evergreen/
2. deciduous/
3. marcescent/
4. withering early/

EVERGREEN: having green leaves through the winter.

DECIDUOUS: falling off, as leaves from a tree; not evergreen; not persistent.

MARCESCENT: having leaves that are withered, dry and possibly brown, but remaining attached to the plant. This may be important in protecting and reducing water loss in plants.

Prophylls of grasses

#61. Prophylls <length>/
mm long/

PROPHYLL: the first leaf of a lateral shoot. It is a leaf sheath with two pronounced veins. Sometimes subsequently produced sheaths without blades may be misinterpreted as prophylls. These may be recognized by having more than two pronounced veins.

#62. Prophylls <features of the veins>/
1. with smooth veins/
2. with scabrous veins/
3. with hairy veins/

PROPHYLL: the first leaf of a lateral shoot. It is a leaf sheath with two pronounced veins.

Sometimes subsequently produced sheaths without blades may be misinterpreted as prophylls. These may be recognized by having more than two pronounced veins.

SMOOTH: devoid of any trichomes (hairs).

SCABROUS: rough to the touch because of small, bristly, prickle-hairs on the surface.

HAIRY: with longer, more slender hairs on the surface.

#63. Prophylls <presence of pronounced keels>/
1. with pronounced keels/
2. lacking pronounced keels/

PROPHYLL: the first leaf of a lateral shoot. It is a leaf sheath with two pronounced veins. Sometimes subsequently produced sheaths without blades may be misinterpreted as prophylls. These may be recognized by having more than two pronounced veins.

KEEL: a raised or pronounced ridge that is V-shaped.

Stipules

#64. Stipules <presence>/
1. present/
2. absent /

STIPULE is one of a pair of appendages at the base of the petiole (the stalk that attaches the leaf to the stem). Stipules are often deciduous, leaving a stipule scar after falling off.

NOTE: To determine if stipules are present check immature leaves (where they should be obvious) or look for stipule scars at the base of mature leaves. A stipule protects a leaf in bud.

#65. Stipules <form>/
1. leaf-like/
2. scale-like/

STIPULE: one of a pair of appendages found at the base of the petiole (the stalk that attaches the leaf to the stem).

SCALE: a thin, usually small and dry structure.

LEAF-LIKE: resembling a leaf.

#66. Stipules <length, measure entire stipule including parts merged with the petiole>/
mm long/

STIPULE: one of a pair of appendages found at the base of the petiole (the stalk that attaches the leaf to the stem).

#67. Stipules <width>/
mm wide/

STIPULE: one of a pair of appendages found at the base of the petiole (the stalk that attaches the leaf to the stem).

#68. Stipules <sheathing>/
1. sheathing <ocreate as in Rumex or connate as in Astragalus>/
2. not sheathing /

STIPULE: one of a pair of appendages found at the base of the petiole (the stalk that attaches the leaf to the stem).

SHEATHING: the stipules form a tube around the stem.

OCREA (=OCREATE): a tubular stipule, or pair of opposite stipules forming a tube.

CONNATE: stipules united at the base in pairs around the petiole.

#69. Stipules <colour>/
1. brown/
2. green/
3. white/
4. colourless/
5. pink or reddish/

STIPULE: one of a pair of appendages found at the base of the petiole (the stalk that attaches the leaf to the stem).

#70. Stipules <surface pubescence>/
1. glabrous/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. woolly/
7. hairs short-silky <less than 0.5 mm>/
8. hairs long-silky <greater than 0.5 mm>/

GLABROUS: without hairs.

PUBESCENT:covered with short, fine hairs.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: bearing short, dense, soft, wooly hairs, that are matted or tangled.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

#71. Stipules <whether glandular>/
1. glandular/
2. without glands /

GLANDULAR: having glands, small bulges that stores or secrete fluids.

#72. Stipules apex <shape>/
1. acuminate/
2. acute/
3. obtuse/
4. rounded/
5. dentate/

STIPULE is one of a pair of appendages at the base of the petiole (the stalk that attaches the leaf to the stem). Stipules are often deciduous, leaving a stipule scar after falling off.

ACUMINATE: gradually tapering to a sharp point and forming concave sides along the tip.

ACUTE: tapering to a point; angle less than 90 degrees, the sides are essentially straight or slightly convex.

OBTUSE: at an angle more than 90 degrees, the sides are essentially straight or slightly convex.

ROUNDED: a rounded apex, approximately 180°

DENTATE: with teeth-like tips.

Petiole

#73. Petioles <presence>/
1. present /
2. absent/

This character refers to leaves on vegetative stems.

PETIOLE: the stalk that attaches the leaf blade to the stem.

#74. Petioles <length>/
mm long/

This character refers to the leaves on the vegetative stems.

PETIOLE: the stalk that attaches the leaf blade to the stem.

NOTE: Leaf characters are based on the largest, mature, medial leaf on the shoot.

#75. Petioles <presence of sessile glands>/
1. with sessile glands/
2. without sessile glands/

GLANDS: small bulges that store or secrete fluids.

#76. Petioles <whether glandular dots at base of leaf>/
1. glandular dots at the base of the leaf present/
2. glandular dots at the base of the leaf absent/

PETIOLE GLANDULAR DOTS: a pair of small glandular lobes or spheres at the base of the leaf.

#77. Petioles <cross-section at midpoint>/
1. convex to flat in cross-section/
2. shallowly concave in cross-section/
3. deeply concave in cross-section, but margins not covering the groove/
4. deeply concave in cross-section, margins covering groove <e.g. some S. reticulata or S. richardsonii>/
5. flat/

PETIOLE: the stalk that attaches the leaf blade to the stem.

NOTE: Leaf characters are based on the largest, mature, medial leaf on the shoot.

Petiole vestiture

#78. Petioles <general hairiness>/
1. glabrous/
2. hairy/
3. glabrescent/
4. scaly/

PETIOLE: the stalk that attaches the leaf blade to the stem.

GLABROUS: without hairs.

GLABRESCENT: initially hairy but becoming glabrous. SCALES: reduced leaf-like structures, several cells wide.

#79. Petioles <surface pubescence type>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. woolly/
7. hairs short-silky <less than 0.5 mm>/
8. hairs long-silky <greater than 0.5 mm>/

PETIOLE: the stalk that attaches the leaf blade to the stem. In Salix specimens this refers to the adaxial surface in particular.

PUBERULENT: minutely pubescent or downy.

PUBESCENT: covered with short, fine hairs.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: with short, dense interwoven hairs.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

NOTE: Leaf characters are based on the largest, mature, medial leaf on the shoot.

#80. Petioles <surface hairs length compared to petiole diameter>/
1. hairs less than the diameter of the petiole/
2. hairs more than the diameter of the petiole/

#81. Petioles hairs <position>/
1. appressed/
2. spreading/
3. erect/
4. reflexed/

APPRESSED: growing at an angle of less than 10 degrees from the culm.

SPREADING: at an angle of 10–80 degrees from the culm.

ERECT: at an angle of 80–90 degrees from the culm.

REFLEXED: the angle is greater than 90 degrees.

#82. Petioles hairs <adaxial hairs shape>/
1. straight/
2. floccose/
3. curved/
4. wavy/
5. crispate /

STRAIGHT: hairs that are not curved, wavy or tufted.

FLOCCOSE: with tufts of soft woolly hairs that often rub off easily. Floccose hairs are flat, irregularly turned, twisted, or felted and appressed to the surface. The hairs are usually so long and intertwined that it is impossile ot tell where a single hair begins or ends. They often form a dense tomentum on the lower leaf surface and petiole but may also occur scattered on the peduncle and inflorescences axes.

CURVED: hairs with a loose curl to 45–120°.

WAVY: with more than one loose curl, like a wave. CRISPTE HAIRS: are unixellular, and less than 1 mm long. The more or less individual hairs are terete, wawy often in a corkscrew-like fashion. They may form a tomentum on the lower leaf surface, or the petiole and sometimes they make the peduncle appear villous. Among our species, such hairs characterise P. pulchella, P. rubricaulis s. lat. and P. vahliana.

#83. Petioles hairs <hair surfaces>/
1. smooth/
2. rough/

Sheaths

#84. Sheaths <longevity>/
1. persisting/
2. breaking down into fibres/

PERSISTING: sheaths not breaking down into fibres.

NOT PERSISTING: the tissue between the vascular bundles disintegrating usually in the first growing season to leave the vascular bundles as fibres.

#85. Sheaths <build up at base of plant>/
1. forming a conspicuous build up at the base of the plant/
2. not forming a conspicuous build up at the base of the plant/

#86. Sheaths <colour>/
1. greyish brown/
2. brown/
3. green/
4. reddish/
5. straw pale yellow/

#87. Sheaths <whether margins are fused>/
1. with the margins fused to the apex/
2. with the margins fused only in the lower part/
3. with the margins not fused/
4. with sheaths fused to the stems and terminating in free, tooth-like leaf tips./

SHEATH: the tubular portion of the leaf, which encloses the stem.

MARGINS FUSED TO APEX: sheath a hollow tube with no overlapping margins at the apex.

MARGINS FUSED ONLY IN THE LOWER PART: margins free and usually overlapping in the upper part.

Most Arctic grasses have state 2. Tropical grasses often have state 3.

#88. Sheaths <surface features>/
1. glabrous/
2. with trichomes/

SHEATH: the tubular portion of the leaf, which encloses the stem.

TRICHOME: any hair or hair-like outgrowth of the epidermis.

#89. Sheaths <type of trichomes>/
1. pubescent/
2. hirsute/
3. scabrous/

SHEATH: the tubular portion of the leaf, which encloses the stem.

PUBESCENT: covered with soft, downy hairs.

HIRSUTE: covered with rather coarse and stiff hairs, these long, straight, and erect or ascending.

SCABROUS: rough to the touch, usually because of the presence of small prickle-hairs on the surface.

BRISTLY: with prickle-hairs.

#90. Collars <monocots, collar, the sheath blade zone, consider adaxial surface>/
1. collars present <sheath abruptly contracted into the blade at a line>/
2. collars absent <sheath tapering into the blade>/

SHEATH: the tubular portion of the leaf, which encloses the stem.

TAPERING INTO THE BLADE: the transition between sheath and blade is gradual.

ABRUPTLY CONTRACTED INTO THE BLADE: The transition between sheath and blade occurs in distinct and a narrow region.

Ligules

#91. Ligules <presence of ligule, Monocot character>/
1. present/
2. absent/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

#92. Ligules <length>/
mm long/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

#93. Ligules <type>/
1. membranous/
2. a fringed membrane/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

MEMBRANOUS : thin, soft, flexible, more or less translucent material, lacking a fringe of trichomes (hairs) on the margin.

FRINGED MEMBRANE: membrane having a fringe of trichomes on the margin.

#94. Ligules <whether hairy>/
1. glabrous/
2. hairy/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

glabrous: without hairs.

#95. Ligules <shape>/
1. linear/
2. lanceolate/
3. ovate-oblong/
4. transversely oblong <long rectangular>/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

OVATE-OBLONG: rectangular in outline, but slightly broader at one end, and attached at the broad end.

TRANSVERSELY OBLONG: in the shape of a rectangle, attached at the long side.

#96. Ligules apices <shape of the apical region>/
1. acuminate/
2. acute/
3. obtuse/
4. truncate/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

ACUMINATE: apical region with sides somewhat concave, tapering to an extended point.

ACUTE: apex formed by margins meeting at less than a 90 degree angle, sides more or less straight.

OBTUSE: at an angle of more than 90 degrees but less than 180 degrees.

TRUNCATE: apical region flat.

#97. Ligules <whether apex entire>/
1. entire/
2. erose/
3. lacerate/
4. cleft/

LIGULE: the membranous projection at the junction of the sheath and blade of the leaf.

ENTIRE: not toothed, notched, or divided, as the continuous margins of some leaves.

EROSE: irregularly and shallowly toothed or gnawed.

LACERATE: irregularly cut or torn.

CLEFT: incised at the midpoint to produce lobes.

#98. <Simple> leaf bases <shape>/
1. cordate/
2. truncate/
3. obtuse/
4. acute/
5. cuneate/
6. attenuate/
7. rounded/
8. hastate/

This character refers to the leaves on the vegetative stems.

BASE: the end of the leaf blade nearest to the point of attachment.

CORDATE: heart-shaped, with the notch at the base.

TRUNCATE: with a squared base, as if it had been cut off.

OBTUSE: at an angle of more than 90 degrees but less than 180 degrees.

ACUTE: tapering to a point with an angle of 45–90 degrees.

CUNEATE: wedge-shaped, triangular; pie-shaped.

ATTENUATE: tapering gradually to a narrow base.

ROUNDED: with a rounded base.

HASTATE: with two pointed lobes projecting at the base.

Juvenile leaves

#99. Juvenile leaves <colour of unfolding leaves>/
1. reddish/
2. yellowish green/

JUVENILE LEAVES: young leaves that are just unfolding.

#100. Juvenile leaves <general hairiness>/
1. glabrous/
2. hairy/

JUVENILE LEAF: young leaf

GLABROUS: without hairs.

#101. Juvenile leaves abaxial <lower> surface <pubescence type>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. woolly/
7. hairs short-silky <less than 0.5 mm>/
8. hairs long-silky <greater than 0.5 mm>/

JUVENILE LEAVES: young leaves that are just unfolding.

PUBERULENT: minutely pubescent or downy.

PUBESCENT: covered with short, fine downy hairs.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but nor matted hairs.

TOMENTOSA: bearing short, dense, interwoven hairs.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

#102. Juvenile leaves hair <density>/
1. sparse/
2. moderately dense <surface 50% visible>/
3. very dense <surface completely obscured>/

JUVENILE LEAVES: young leaves that are just unfolding.

#103. Juvenile leaves hair <colour>/
1. white/
2. grey/

JUVENILE LEAVES: young leaves that are just unfolding.

Blade measurements

#104. <Simple or compound leaf> blades <length excluding petiole>/
mm long/

This character refers to the leaves on the vegetative stems. The length of the leaf blade is measured from the tip of the leaf blade to the top of the petiole.

PETIOLE: the stalk that attaches the leaf blade to the stem.

NOTE: Leaf characters are based on the largest, mature leaf on a shoot.

#105. <Simple or compound leaf> blades <total width>/
mm wide/

This character refers to the leaves on the vegetative stems. The width of the leaf blade is measured at the widest point of the leaf blade. Width is measured as seen to the naked eye: actual width on a flat leaf, approximately half the true width of a folded leaf. With compound leaves the width is measured between the tips of adjacent leaflets in the middle of the leaf.

#106. <Leaf or leaflet> blades length-width ratio/

To measure leaf shape determine the leaf length and width, then calculate the length/width ratios.

Leaf blades

#107. <Leaf or leaflet> blades <orientation>/
1. appressed to the stem/
2. spreading from the vertical/
3. divaricate/
4. reflexed/

APPRESSED: growing at an angle of less than 10 degrees from the culm.

SPREADING: at an angle of 10–80 degrees from the culm.

DIVARICATE: at an angle of 80–90 degrees from the culm.

REFLEXED: the angle is greater than 90 degrees.

#108. <Leaf or leaflet> blades <texture in fresh plants>/
1. herbaceous /
2. leathery/
3. succulent/
4. membranous/

Character may be difficult to interpret in dried material.

HERBACEOUS: thin, soft, flexible and usually green.

LEATHERY: leather-like.

SUCCULENT: flesh-like; plump, pulpy.

MEMBRANOUS: thin, soft flexible, and more or less translucent, like a membrane.

#109. <Leaf or leaflet> blades <appearance>/
1. straight/
2. somewhat curled/
3. circinate when young/

SOMEWHAT CURLED: leaves curved or curled away from vertical.

CIRCINATE: coiled from the top downwards, into a ring or partially so.

#110. <Leaf or leaflet> blades <usually ferns or monocots, vernation>/
1. rolled in bud/
2. folded in bud/

ROLLED IN BUD: leaf margins overlapping.

FOLDED IN BUD: leaf margins touching.

#111. <Simple leaf> blades <shape>/
1. linear/
2. oblong/
3. elliptic/
4. circular/
5. lanceolate/
6. ovate/
7. oblanceolate/
8. obovate/
9. spatulate/
10. reniform/
11. triangular/
12. lyrate/

This character refers to the leaves on the vegetative stems.

NOTE: When using leaf shape, choose leaves from the lower part of the stem.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends.

CIRCULAR: approximately circular in outline.

LANCEOLATE: lance or spear-shaped; much longer than wide, with the widest point below the middle.

OVATE: egg-shaped in outline and attached at the broad end.

OBLANCEOLATE: inversely lanceolate, with the widest point above the middle.

OBOVATE: inversely ovate, with the attachment at the narrower end.

SPATULATE: spoon-shaped, with a rounded blade above gradually tapering to the base.

RENIFORM: kidney shaped.

TRIANGULAR OR DELTOID: shaped like an equilateral or Isosceles triangle.

LYRATE: a pinnatifid leaf in which the terminal lobe is large and rounded and the lower and lateral lobes are smaller.

#112. <Leaf> blades <Monocots, presence of blade or sheath auricles>/
1. with sheath auricles/
2. with blade auricles/
3. without auricles /

AURICLE: an appendage of various shapes, often ear shaped, found at the junction of the leaf blade and sheath.

BLADE AURICLES: are found at the base of a leaf blade and usually at right angles to it.

SHEATH AURRICLES: are at the apex of the sheath and usually erect and parallet with the blade..

#113. <Simple leaf> blades <shape in cross section; mainly for monocots>/
1. circular in cross section/
2. triangular in cross section/
3. flat/
4. bristle-like/
5. strongly keeled/
6. involute/
7. revolute/
8. folded/
9. caniculate/
10. convolute/

CIRCULAR: outline of a transverse section of the leaf is a circle.

TRIANGULAR: outline of a transverse section of the leaf appears like an equilateral triangle, sometimes with rounded corners. FLAT: the transverse section in one plane.

BRISTLE-LIKE: thin and stiff like a bristle.

INVOLUTE: with margins rolled inwards.

REVOLUTE: with margins rolled outwards.

STRONGLY KEELED: with a prominent midvein.

FOLDED: leaves V-shaped or loosely curved in cross section, margins more or less parallel to each other.

FOLDED or CONDUPLICATE: folded so that the sides face each other.

CANICULATE: leaves essentially circular in cross-section with a groove running down the adaxial surface.

CONVOLUTE: overlapping with one edge and curled inwards.

Leaf blade venation

#114. <Leaf or leaflet> blades <venation pattern>/
1. with pinnate veins/
2. with three main veins <trinerved>/
3. with palmate veins/
4. with parallel veins/
5. appearing single-veined/
6. with inconspicuous veins/
7. with reticulate veins/

VENATION: the pattern of veining on a leaf blade, usually most easily seen on the lower surface of the leaf.

PINNATE: a pattern of venation with veins arranged on opposite sides of one main vein.

TRINERVED: appearing to have three nerves arising from near the base.

PALMATE: a pattern of venation with veins radiating from a common point, like the fingers on a hand.

PARALLEL: a pattern of venation with main veins parallel to the leaf axis or to each other.

APPEARING ONE-VEINED: a pattern of venation where one vein is conspicuous; other veins may be present but they are not easily seen.

RETICULATE: veins in a network pattern.

#115. <Leaf or leaflet> blades <grasses, midvein size>/
1. midvein conspicuously larger than the lateral veins/
2. midvein similar in size to other veins in the leaf/

MIDVEIN: the central vein (nerve) of a leaf.

#116. <Leaf or leaflet> blades <usually grasses, presence or absence of bulliform cells associated with the midvein>/
1. bulliform cells in distinct rows on either side of the midvein/
2. without bulliform cells in a distinct row on either side of the midvein /
3. bulliform cells in a single row above the midvein resulting in the leaf looking longitudinally channelled/

BULLIFORM CELLS: usually large, thin-walled, highly vacuolated, colourless epidermal cells present in the intercostal zones of the leaf blade. These are most commonly present at the base of furrows on the adaxial surface, but may also be present on the abaxial surface. They may be visible as clear lines down either side of the midvein.

#117. <Leaf or leaflet> blades <whether septate nodulose>/
1. septate nodulose/
2. not septate nodulose /

SEPTATE NODULOSE: having raised, prominent end walls to cells.

#118. <Leaf or leaflet> blades secondary veins <whether protruding>/
1. impressed into adaxial <upper> surface, protruding on abaxial <lower> surface/
2. protruding on adaxial <upper> and abaxial <lower> surfaces/
3. flat on adaxial <upper> surface, protruding on abaxial <lower> surface/

SECONDARY VEINS: are veins arising from the midrib.

IMPRESSED: leaf veins situated below the surface; usually the leaf has a thicker mesophyll. MESOPHYLL: the central tissues of a leaf between the upper and lower epidermis.

PROTRUDE: leaf veins that partially stick out.

FLAT: leaf veins situated at the surface; usually the leaf has a thin mesophyll.

NOTE: The apparent position of the secondary veins can intergrade and may be an artefact of a dried specimen.

#119. <Leaf or leaflet> blades secondary veins <position>/
1. arising along midrib/
2. mostly arising at or close to a single point at base and running in recurved arches toward leaf apex/

MID-RIB: the main vein of a leaf.

SECONDARY VEINS: are veins arising from the midrib.

#120. <Leaf or leaflet> blades <stomatal patterns on largest mature leaves>/
1. stomata only on abaxial <lower> surface/
2. stomata on both adaxial <upper> and abaxial <lower> surfaces/
3. stomata on adaxial <upper> surface only present along veins or at apex/

STOMATA: a pore in the plant epidermis, through which gaseous exchange occurs.

NOTE: Stomata can appear as minute, pale dots under about 10–25X magnification and sometimes to the naked eye on the upper surface.

Blade vestiture

#121. <Leaf or leaflet> blades <presence of hairs> /
1. glabrous/
2. scabrous/
3. hairy/

This character refers to the leaf blade and not the leaf margin.

GLABROUS: without hairs.

SCABROUS: rough to the touch, usually because of the presence of small prickle-hairs on the surface.

#122. <Leaf or leaflet> blades <location of hairs on surfaces, excluding margins>/
1. with hairs on the adaxial <upper> surface only/
2. with hairs on the abaxial <lower> surface only/
3. with hairs equally dense on both surfaces/
4. with hairs of different densities on both surfaces/

Adaxial (upper) blade surface

#123. <Leaf or leaflet> blades adaxial <upper> surface <luster>/
1. dull /
2. shiny/
3. highly glossy/
4. glaucous/

DULL: with a surface similar to that of flat paint.

SHINY: with a surface similar to semi-gloss paint.

HIGHLY GLOSSY: with a surface similar to high gloss paint.

GLAUCOUS: covered with a whitish or bluish waxy coating (bloom), as on the surface of a plum.

#124. <Leaf or leaflet> blades adaxial <upper> surface <features>/
1. glabrous/
2. glabrescent <becoming glabrous, usually sparsely hairy>/
3. scabrous/
4. hairy/

ADAXIALLY: on the side facing the main axis of the plant, i.e. the upper surface of the leaf.

glabrous: without hairs.

GLABRESCENT: initially hairy but becoming glabrous.

SCABROUS: rough to the touch because of small, bristly, prickle-hairs.

HAIRY: with longer hairs.

#125. <Leaf or leaflet> blades adaxial <upper> surface hairs <pubescence type>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. short-silky <hairs less than 0.5 mm>/
6. long-silky <hairs greater than 0.5 mm>/
7. strigose/

PUBERULENT: minutely pubescent or downy.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

SILKY: straight, appressed, shiny hairs.

STRIGOSE: covered with straight, stiff hairs or bristles.

PUBESCENT: covered with short, fine hairs easilly visible to the naked eye.

#126. <Leaf or leaflet> blades adaxial <upper> surface hairs <density>/
1. sparse/
2. moderately dense <surface 50% visible>/
3. dense/

#127. <Leaf or leaflet> blades adaxial <upper> surface hairs <hair colour>/
1. white and translucent/
2. grey/
3. a mixture of white and rust-coloured/
4. a mixture of white and yellow/
5. multicellular with deep purple pigment at the cell end walls/

#128. <Leaf or leaflet> hairs on blades <hair type>/
1. simple, unbranched without glands/
2. branched/
3. glandular/
4. stellate/

GLANDULAR: bearing glands, small bulges that secrete sticky or oily substances.

STELLATE: hairs arranged in the form of stars, with several to many branches radiating from the base.

BRANCHED: with two or more forking divisions.

#129. <Leaf or leaflet> blades adaxial <upper> surface <sessile glands> /
1. with sessile glands/
2. without sessile glands/

SESSILE GLANDS: small bulges on the surface that store or secrete fluids.

Abaxial (lower) blade surface

#130. <Leaf or leaflet> blades abaxial <lower> surface <luster>/
1. dull /
2. shiny/
3. highly glossy/
4. glaucous/

DULL: with a surface similar to that of flat paint.

SHINY: with a surface similar to semi-gloss paint.

HIGHLY GLOSSY: with a surface similar to high gloss paint.

GLAUCOUS: covered with a whitish or bluish waxy coating (bloom), as on the surface of a plum.

#131. <Leaf or leaflet> blades abaxial <lower> surface <glaucescence or waxy bloom>/
1. not glaucous/
2. glaucous/

GLAUCOUS: covered with a whitish or bluish waxy coating (bloom), as on the surface of a plum.

#132. <Leaf or leaflet> blades abaxial <lower> surface <sessile glands>/
1. with sessile glands/
2. without sessile glands/
3. glands (if present) not visible/
4. with glandular hairs/

SESSILE GLANDS: small bulges on the surface that store or secrete fluids.

#133. <Leaf or leaflet> blades abaxial <lower> surface <hairiness>/
1. glabrous/
2. glabrescent/
3. scabrous/
4. hairy/
5. scaly/

ABAXIALLY: on the side facing away from the main axis of the plant, i.e. the underside of the leaf.

GLABROUS: without hairs.

GLABRESCENT: initially hairy but becoming glabrous.

SCABROUS: rough to the touch because of small, bristly, prickle-hairs.

HAIRY: with longer, more slender hairs on the surface.

SCALY: small flakes like fish scales.

#134. <Leaf or leaflet> blades abaxial <lower> surface <pubescence type>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. hairs short-silky <less than 0.5 mm>/
7. hairs long-silky <greater than 0.5 mm>/
8. hairs woolly/
9. strigose/

PUBESCENT: covered with short, fine hairs easilly visible to the naked eye.

PUBERULENT: covered with short, fine hairs, better seen at 10x or 40x magnification.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: with short, dense interwoven hairs.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

STRIGOSE: covered with straight, stiff hairs or bristles.

#135. <Leaf or leaflet> blades abaxial <lower> surface hairs <density>/
1. sparse/
2. moderately dense/
3. very dense/

SPARSE: surface mostly visible.

MODERATELY DENSE: surface about 50% visible.

DENSE: surface completely obscured.

#136. <Leaf or leaflet> blades abaxial <lower> surface hairs <hair colour>/
1. white, or translucent/
2. rust-coloured/
3. amixture of white, or translucent, and rust-coloured/
4. a mixture of white and yellow/

#137. <Leaf or leaflet> blades abaxial <lower> surface hairs <shape>/
1. straight/
2. curved/
3. wavy/

STRAIGHT: hairs that are not curved, wavy or tufted.

CURVED: hairs with a loose curl to 45–120°.

WAVY: with more than one loose curl, like a wave.

#138. <Leaf or leaflet> blades abaxial <lower> surface hairs <hair orientation>/
1. appressed/
2. spreading/
3. erect/

APPRESSED: hair pressed close or flat against the branchlet.

SPREADING: hair standing out from the surface.

ERECT: hair vertical, not declining or spreading.

Blade margins

#139. <Simple> leaves <whether lobed>/
1. lobed/
2. not lobed /
3. cut into linear divisions/

LOBED: bearing lobes cut less than half way to the base.

LINEAR DIVISIONS: leaf divided into lobes that are cut more than half way to the base.

#140. <Simple or compound> leaf margins <toothing>/
1. entire /
2. glandular-dotted/
3. serrulate/
4. serrate/
5. crenate/
6. dentate/
7. deeply divided/

MARGIN: the edge of the leaf blade.

ENTIRE: not toothed; the margins smooth.

GLANDULAR-DOTTED: margins are entire but there are glandular dots, small bulges that secrete fluid on the edge.

SERRULATE: minute, sharp teeth pointing forward.

CRENATE: rounded teeth, scalloped along the margin.

SERRATE: sharp large teeth pointing forward.

DENTATE: or toothed, a leaf margin with sharp teeth or indentations pointing outwards at right angles to the midrib.

DEEPLY DIVIDED: Leaf blade with divisions cut more than half way to the base; these may someteimes be mistaken for compound leaves as in Myriophyllum.

#141. <Simple or compound> leaf margins with teeth <position>/
1. all around leaf/
2. toward base only/
3. toward apex only/

MARGIN refers to the edge of the leaf blade.

TEETH: sharp pointed strutures like a saw or sharks teeth.

#142. <Simple or compound> leaf margins with <number of teeth> teeth per cm/

MARGIN refers to the edge of the leaf blade.

TEETH: sharp pointed strutures like a saw or sharks teeth.

#143. <Leaf or leaflet> leaf margins, degree of incision <as a percentage. previously, 1. less than 10% 2. 20–25% 3. 25–50% 4. 50–75% 5. more than 75%>/
%/

#144. <Leaf or leaflet> leaf margins, with <number of> teeth on one side of blade <of a leaf or apical leaflet in compound leaves; previously, 1. 1–2 2. 3–5 3. 6–10 4. more than 10>/

TEETH: sharp pointed strutures like a saw or sharks teeth.

#145. <Simple or compound> leaf margins <glands position>/
1. with marginal glands/
2. with submarginal glands/

MARGIN refers to the edge of the leaf blade.

SUBMARGINAL: glands borne on the upper surface, slightly in from the edge. GLANDS; small bulges that store or secrete fluids.

#146. <Simple or compound> leaf margins with sessile glands <position>/
1. all around leaf/
2. toward base only/
3. toward apex only/

SESSILE GLALNDS. small bulges that store or secrete fluids.

#147. <Simple or compound> leaf margins with <number of glands per cm>/
glands per cm/

#148. <Simple or compound> leaf margins <whether hairy>/
1. glabrous /
2. scabrous/
3. with non-glandular hairs/
4. with glandular hairs/

GLABROUS: without hairs or scarbrous trichomes.

SCABROUS: with stiff hairs that contain silica and feel prickly to the touch.

NON-GRANDULAR: with soft textured usually straight hairs without silica, or globular end glands.

GLANDULAR HAIRS: hairs that have often globular ends that store or secrete fluids.

#149. <Simple or compound> leaf margins with glandular hairs <position>/
1. all around leaf/
2. toward base only/
3. toward apex only/

GLANDULAR HAIRS: hairs that have often globular ends that store or secrete fluits.

#150. Conspicuous hydathodes <presence>/
1. present/
2. absent /

HYDATHODES: specialized pores (small openings) at the end of veins that exude water and dissolved minerals which may be seen as white lime deposits.

Leaf blade apices

#151. <Simple or compound> leaf apices <tip shape, tip of uppermost leaflet in a compound leaf>/
1. acuminate/
2. acute/
3. obtuse/
4. rounded/
5. retuse/
6. mucronate/

This character refers to the leaves on the vegetative stems.

APEX, (plural APICES): the tip; the farthest point of attachment of the leaf to the stem.

ACUMINATE: gradually tapering to a sharp point at an angle of less than 45 degrees usually forming concave sides along the tip.

ACUTE: tapering to a point with an angle of 45–90 degrees.

OBTUSE: at an angle of more than 90 degrees but less than 180 degrees.

ROUNDED: a rounded apex.

RETUSE: with a shallow notch in a round or blunt apex.

MUCRONATE: with a sharp point, short spur, or spiny tip.

Compound leaves

#152. <Compound> leaflet arrangement <form>/
1. trifoliate/
2. pinnate/
3. digitate/
4. decompound/

TRIFOLIATE: a compound leaf which is divided into three leaflets.

PINNATE: a compound leaf with leaflets arranged in two rows, one on each side of the midrib.

DIGITATE: a compound leaf with 4 or more leaflets divided from a common point, like the fingers of a hand.

DECOMPOUND: leaves that are more than once divided, or compound.

#153. Leaflets <number>/

LEAFLET: a division of a compound leaf.

#154. Leaflets <length in mm>/
mm long/

LEAFLET: a division of a compound leaf. Measure length of the terminal leaflet; in the legumes measure leaflet nearest the middle of the blade.

#155. Leaflet <width in mm>/
mm wide/

LEAFLET: a division of a compound leaf. Measure the width of the terminal leaflet; in legumes, measure leaflet nearest the middle of the blade.

#156. Leaflets <shape>/
1. linear/
2. oblong/
3. elliptic/
4. ovate/
5. obovate/
6. lanceolate/
7. oblanceolate/
8. lacinate/
9. triangular/
10. obtriangular/

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends.

OVATE: egg-shaped in outline and attached at the broad end.

OBOVATE: inversely ovate, with the attachment at the narrow end.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

OBLANCEOLATE: inversely lanceolate, with the widest point above the middle.

LACINATE: divided into several long and more or less equal segments.

TRIANGULAR: having the shape of a triangle with two equal side and the apex pointing up.

OBTRIANGULAR:having the shape of a triangle with two equal side and a flat surface pointing up.

#157. Leaflet <whether veins conspicuous>/
1. veins conspicuous/
2. veins inconspicuous/

VEIN: general term for the plant plumbing made up of xylem that carries the water and phloem that carries sugars and associated tissues.

#158. <Compound> leaflet base <whether the apical leaflet is stipitate>/
1. of apical leaflet distinctly stipitate/
2. of apical leaflet not distinctly stipitate /

APICAL LEAFLET: in a compound leaf that has an odd number of leaflets, this is the terminal unmatched leaflet. STIPITATE: having a stalk or stipe.

#159. <Compound> leaflet base <length of apical leaflet> stipe/
mm long/

STIPE: the midvien of the leaf: here, the short stalk between uppermost leaflets and the apical leaflet.

#160. Plants <sexuality>/
1. monoecious/
2. dioecious/
3. bisexual /
4. asexual/
5. reproducing by spores borne in sporangia/

MONOECIOUS: having both male and female reproductive organs on the same plant but separated in different floral structures. Having unisexual flowers of different sexes on the same plant.

DIOECIOUS: having the female and male flowers on different plants.

ASEXUAL: reproducing without sexual union.

BISEXUAL: having both male and female reproductive organs in the same flower or capitulum.

SPORANGIA(plural): SPORANGIUM (singular), a case containing spores that develop into a tiny plant, the gametophyte generation. Character that separates the Pteriodphytes. In the Arctic these are in the families Equisetaceae, Lycopodiaceae and Polypodiaceae.

Pteriodophytes leaves

#161. Sporangia <position>/
1. in the axiles of unmodified leaves/
2. in terminal cone-like structures/
3. in sori on the under-surface of the leaves/

SPORANGIA: (plural) SPORANGIUM: (singular): a case containing spores that develop into a gametophte (gamete producing) generation of the plant. These plants are tiny and rarely found.

#162. Sori <presence of an indusium on sori>/
1. with a distinct indusium/
2. without a distinct indusium/

SORI: (plural): SORUS (singular): a heap or cluser of sporangia.

INDUSIUM: the covering over a sporangium that may be a flap of tissue or in Woodsia Numerous multicellular hairs.

#163. Indusia <structure>/
1. of narrow hair-like segments, one row of cells many times longer than wide, and longer than the sporangia/
2. ovate, whitish, lanceolate scales that fall early/

Flowering stems (peduncles)

#164. Flowering stems <or culms, length relative to leaves>/
1. shorter than the leaves/
2. about as high as the leaves/
3. conspicuously taller than the leaves/

#165. Flowering stems <peduncle> <length from base to lowest flower, or inflorescence branching>/
cm long/

PEDUNCLE: or flowering stem, a primary flower stalk supporting either a cluster of flowers, a capitulum, or a solitary flower, the stalk above a leaf subtending the inflorescence. In Juncaceae the stalk above the uppermost leaf, or reduced leaf subtending the inflorescence.

#166. Flowering stems <peduncles, presence of leaves>/
1. with leaves <sometimes smaller than basal leaves>/
2. without leaves <scapose>/
3. without leaves in the upper half of the peduncle/

FLOWERING STEM (Peduncle): an elongated stem, usually arising from a mostly basal cluster of leaves, having a terminal flower or inflorescence. It may, or may not, have leaves or bracts.

#167. Flowering stems uppermost leaf <position on the flowering stems>/
1. arising below the middle of the stem/
2. arising above the middle of the stem/

This character was developed for the Cyperaceae, it may not be appropriate in other families.

#168. Flowering stems <peduncle> <hairs>/
1. glabrous/
2. hairy/
3. with sessile glands/

FLOWERING STEM: an elongated stem, usually arising from a mostly basal cluster of leaves, having a terminal flower or inflorescence. It may, or may not, have leaves or bracts.

#169. Flowering stem <peduncle surface pubescence>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. short-silky <hairs less than 0.5 mm>/
7. long-silky <hairs greater than 0.5 mm>/
8. woolly/

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: with short, dense interwoven hairs.

SILKY: straight, appressed, shiny hairs.

PUBESCENT: covered with short, fine hairs easilly visible to the naked eye.

PUBERULENT: covered with short, fine hairs, better seen at 10x or 40x magnification.

FLOCCOSE:

#170. Flowering stems <peduncle> hairs <type>/
1. simple <unbranched>/
2. branched/
3. glandular/
4. stellate/

#171. Flowering stem hairs <length compared to peduncle diameter>/
1. less than the diameter of the peduncle/
2. more than the width of the peduncle/

#172. Flowering stems <peduncle> hairs <colour>/
1. white or translucent/
2. brown/
3. black/
4. transparent with red cross-walls/
5. yellow/
6. transparent with deep purple cross-walls/

FLOWERING STEM: peduncle: the stalk of a single flower, or group of flowers. PEDICEL: the stalk of a single flower in a group of flowers.

#173. Flowering culm nodes <whether rooting at the lower culm nodes, grasses>/
1. rooting at the lower nodes/
2. not rooting at the lower nodes /

NODE: the area on a stem (culm) where one or more leaves are borne.

Culms of grasses

#174. Flowering culm nodes <whether exposed>/
1. not exposed/
2. becoming exposed/

NODE: the area on a stem (culm) where one or more leaves are borne.

NOT EXPOSED: sheaths longer than the internode.

EXPOSED: sheaths shorter than the internode.

In the Arctic, the plants frequently do not elongate enough for nodes to be visible. In a limited number of species, however, whether or not nodes are visible can be a useful indication of whether sheaths are longer or shorter than the internodes.

#175. Flowering culm nodes number visible <if exposed, applies to grasses>/

NODE: the area on a stem (culm) where one or more leaves are borne.

NOT EXPOSED: sheaths longer than the internode.

EXPOSED: sheaths shorter than the internode.

In the Arctic, the plants frequently do not elongate enough for nodes to be visible. In a limited number of species, however, whether or not nodes are visible can be a useful indication of whether sheaths are longer or shorter than the internodes.

#176. Flag leaf sheaths <whether inflated>/
1. inflated/
2. not inflated /

FLAG LEAF: uppermost culm leaf of grasses.

INFLATED: puffed out, loosely fitting around the stem.

Leaves or bracts on sedge culms

#177. Leaf or reduced bract closely associated with the base of the inflorescence <presence>/
1. present/
2. absent/

REDUCED BRACT: a leaf-like or scale-like structure in sedges, similar to a flag leaf in grasses. This is a sedge character that does not apply to the Asteraceae.

#178. Leaf or reduced bract closely associated with the base of the inflorescence <whether conspicuous>/
1. conspicuous and leaf-like/
2. reduced, or scale-like/

#179. Leaf or reduced bract closely associated with the base of the inflorescence <relative length>/
1. exceeding the inflorescence/
2. similar in length to the inflorescence/
3. shorter than the apex of the inflorescence/

INFLORESCENCE: is a cluster of flowers.

#180. Leaf or reduced bract closely associated with the base of the inflorescence <length>/
mm long/

Measured from leaf tip to the point where the leaf is appressed to the stem.

#181. Leaf or reduced bract closely associated with the base of the inflorescence <relative sheath length>/
1. with sheath longer than the blade/
2. with sheath shorter than the blade/
3. sheathless/

SHEATH: the tubular portion of the leaf, which encloses the stem.

BLADE: the flat expanded portion of a leaf.

#182. Leaf or reduced bract closely associated with the base of the inflorescence <longevity>/
1. caducous/
2. persistent /

CAUDUCOUS: falling early before the fruit is mature.

#183. Leaf or reduced bract closely associated with the base of the inflorescence <whether tip is calloused>/
1. with calloused tip/
2. without calloused tip /

Inflorescence

#184. Flowers <grouping>/
1. solitary/
2. in inflorescences /

SOLITARY: one flower; not in clusters; borne singly.

INFLORESCENCE: a cluster of flowers that are rarely replaced by vegetative bulblets.

Not to be recorded for Asteraceaea.

#185. Inflorescences <type>/
1. spicate/
2. racemose/
3. paniculate/
4. cymose/
5. fasciculate/
6. head-like/
7. a catkin/
8. an umbel/
9. a solitary capitulum <or head>/
10. of several flowering heads (capitulesence)/
11. a dichasium/

INFLORESCENCE: a cluster of flowers that are rarely replaced by vegetative bulblets. In grasses, the flowering portion of the culm, delimited at the base by the uppermost leafy node of the shoot.

SPICATE: an inflorescence with sessile or subsessile flowers or spikelets attached directly to the main axis. Flowers or spikelets maturing from the bottom upwards.

RACEMOSE: an inflorescence with pedicelled flowers or spikelets attached directly to the main axis and maturing from the bottom upwards.

PEDICEL: the stalk of an individual flower in an inflorescence or the stalk of a grass spikelet or sedge secondary spike.

PANICULATE: a branched inflorescence with pedicelled flowers, maturing from the bottom upwards.

CYMOSE: a branched flat-topped or round-topped inflorescence, with the terminal flowers blooming first,

FASCICULATE: a tight bundle or clusters of pedicelled flowers originating from a common point.

HEAD-LIKE: a dense cluster of sessile (no pedicel) or subsessile (slight pedicel) flowers, sometimes with as few as two flowers as in Juncus biglumis

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous (no petals), unisexual flowers.

UMBEL: an inflorescence in which the stalks of the flowers all arise from the top of the main stalk.

CAPITULUM OR HEAD (plural capitula): a dense inflorescence usually comprised of sessile small flowers that appear to be a single flower (dandelion).

DICHASIUM: inflorescence in which two lateral branches occur at about the same level with flowers at approximately the same stage of development.

CAPITULESENCE: an inflorescence composed of more than one capitulum.

#186. Inflorescence <or single flower> <position>/
1. terminal /
2. axillary/

INFLORESCENCE: a cluster of flowers that are rarely replaced by vegetative bulblets.

TERMINAL: located at the end of the stem.

AXILLARY: arising in the leaf axil between the stem and the leaf.

#187. Inflorescence <whether dense>/
1. dense/
2. diffuse/

INFLORESCENCE: the flowering portion of the culm, delimited at the base by the uppermost leafy node of the shoot.

DENSE: Flowers or spikelets tightly grouped.

DIFFUSE: with the branches spread out, so that the inflorescence is not dense. This character should be assessed when the plant is at or after anthesis; most inflorescences will likely be compact and considered dense before this time.

#188. Inflorescence <shape>/
1. linear/
2. oblong/
3. lanceolate/
4. ovate/
5. globose or subglobose/
6. obovate/
7. ellipsoid/
8. cylindrical/

INFLORESCENCE: the flowering portion of the culm, delimited at the base by the uppermost leafy node of the shoot.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

OVATE: egg-shaped in outline and attached at the broad end.

GLOBOSE: shaped like a round ball.

OBOVATE: inversely ovate, with the attachment at the narrow end.

ELLIPSOID: football-shaped; a three dimensional structure that is elliptic in long section and circular in cross section.

CYLINDRICAL: cylinder-shaped, elongate and round in cross-section.

Character is not to be scored for diffuse inflorescences.

#189. Inflorescence <total length including all separate spikes>/
cm long/

INFLORESCENCE: the flowering portion of the culm, delimited at the base by the uppermost leafy node of the shoot.

In grasses the length is measured from the bottom node to the top of the inflorescence, excluding the awns.

#190. Inflorescence <width>/
mm wide/

INFLORESCENCE: the flowering portion of the culm, delimited at the base by the uppermost leafy node of the shoot.

The width is measured at the widest point of the inflorescence, and includes the awns.

#191. Inflorescence <whether elongate in fruit>/
1. not elongating as the fruit matures/
2. elongating as the fruit matures/

ELONGATING AS FRUIT MATURES:

#192. Inflorescence <bulbils>/
1. with bulbils/
2. without bulbils /

INFLORESCENCE: a cluster of flowers that are rarely replaced by vegetative bulbils.

BULBIL: a small bulb borne in a leaf or bract axil (point between the stem or bract and a leaf). They can easily detach and fall to the ground and vegetatively propagate a new plant (a form of vegetative propagation).

Inflorescence main axis and branches in grasses

#193. Inflorescence main axis <rachis> <hairs>/
1. glabrous/
2. scabrous/
3. hairy/

INFLORESCENCE: the flowering portion of the culm, delimited at the base by the uppermost leafy node of the shoot.

Glabrous: without hairs.

SCABROUS: rough to the touch because of small, bristly, prickle-hairs.

HAIRY: with longer hairs.

These character states are distinguishable with the naked eye or at 10X magnification.

RACHIS: stalk between the flowers of an inflorescence.

#194. Inflorescence main branches angle of divergence/
1. less than 30 degrees/
2. 30–60 degrees/
3. 60–90 degrees/
4. more than 90 degrees/

Measure the angle between the rachis and the upper surface of the branch.

#195. Number of inflorescence branches at lowest node/

NODE: area on a stem bearing one or more leaves.

#196. Inflorescence primary branches <measure longest at lowest node, length>/
mm long/

The length of the longest branch on the plant, to the base of the terminal spikelet.

#197. Inflorescence primary branches <surface features>/
1. glabrous/
2. scabrous/

GLABROUS:

SCABROUS: with small, prickle-hairs.

#198. Inflorescence primary branches <whether the secondary branches are spreading>/
1. with appressed secondary branches/
2. with spreading secondary branches/

APPRESSED: branches growing at an angle of less than 10 degrees from the rachis or main stem.

Pedicels

#199. Pedicels <surface>/
1. glabrous/
2. scabrous/
3. hairy/
4. with glandular hairs/
5. with sessile glands/

PEDICEL: the stalk of an individual flower in an inflorescence or the stalk of a grass spikelet or sedge secondary spike.

SMOOTH: without any roughness.

SCABROUS: rough to the touch because of small, bristly, prickle-hairs.

PUBESCENT: covered with short, fine hairs, downy.

GLANDULAR HAIRS: capable of secreting products, frequently stalked.

#200. Pedicels bract leaves <length>/
mm long/

PEDICEL: the stalk of an individual flower in an inflorescence or the stalk of a grass spikelet or sedge secondary spike.

BRACT LEAVES: reduced leaves on a flowering stem or pedicel, usually, but not always closely associated with a flower or flower cluster.

#201. Pedicels bract leaves <width>/
mm wide/

PEDICEL BRACT LEAVES: small leaves on the pedicel not immediately adjacent to the flower. Character used in the Caryophyllaceae.

#202. Pedicels bract leaves <margins whether scarious>/
1. with a distinct scarious margin/
2. without a distinct scarious margin/

SCARIOUS MARGIN: a thin, dry, non-green often translucent edge. Character used in the Caryophyllaceae.

#203. Pedicels bract leaves <width of scarious margin>/
1. margins less than 0.3 mm/
2. margins 0.3–0.8 mm/

SCARIOUS MARGIN: a thin, dry, non-green often translucent edge.Character used in the Caryophyllaceae.

Spikes of sedges

#204. Inflorescence <whether uni- or multispicate>/
1. a single spike/
2. multispicate/

SPIKE: an unbranched, indeterminate, elongated inflorescence which bears sessile flowers.

MULTISPICATE: composed of two or more spikes.

#205. Inflorescence <total number of spikes in a multispicate inflorescence>/
spikes/

#206. Inflorescence <whether lateral spikes are sessile>/
1. lateral spikes sessile/
2. lateral spikes borne on pedicels/

PEDICEL: the stalk of an individual flower in an inflorescence or the stalk of a grass spikelet or sedge secondary spike.

SESSILE: the catkin is attached directly onto the stem.

#207. Individual spike(s) <orientation>/
1. erect/
2. ascending <but not erect>/
3. divergent <but neither erect nor ascending>/
4. reflexed/
5. pendent <drooping>/

ERECT: stiffly vertical.

ASCENDING: within the range of angles between erect and divergent.

DIVERGENT: more of less at right angles to the inflorescence.

REFLEXED: bent back, rigidly turned over, and maybe downwards.

PENDENT: hanging or drooping down.

#208. Bisexual spike(s) <Eriophorum, presence of floral bracts at the base>/
1. with empty bracts at the base/
2. without empty bracts at the base /

BRACTS: scale-like structures immediately below a flower as in Eriophorum or Kobresia.

#209. Terminal spike <sexuality>/
1. staminate at the base/
2. wholly staminate/
3. staminate at the apex/
4. with both sexes in each floret <bisexual>/
5. pistillate/

TERMINAL spike, the uppermost spike on the flowering stem.

STAMINATE: male flowers with anthers.

PISTIL: either a single carpel (female reproductive organs) or a group of fused carpels

PISTILLATE: female flowers with ovaries.

#210. Cladoprophyll <presence in Carex>/
1. present at the base of the peduncle of lateral spikes <characteristic of subgenus Carex>/
2. absent <characteristic of subgenus Vignea>/

CLADOPROPHYLL: an inner sheath, a tiny hyaline (usually) tubular prophyll that wraps around the peduncle of a Carex spike.

#211. Capitulum <types of florets present>/
1. with only disc florets/
2. with only ray <ligulate> florets/
3. with ray and disc florets/

RAY FLORET: usually, assymetrical, (ligulate) floret with fused petals open on one side almost to wth base e.g. in Asteraceae head.

DISC FLORET: symmetrical floret at the centre of an Asteraceae head.

Involucres

#212. Involucral bracts <inflorescence whether involucral bracts, phyllaries are present>/
1. present/
2. absent /
3. flowers on pedicels that have bracts leaves/

INFLORESCENCE: a cluster of flowers that are rarely replaced by vegetative bulbils.

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families.

BRACT: a much-reduced leaf, particularly the small or scale-like leaves in a flower cluster or associated with the flowers.

Pedicels that have bracts leaves at some distance from the flower are found in the Caryophyllaceae.

#213. Involucral bracts <shape>/
1. with a flat blade surface/
2. with blades cut into linear divisions/

INVOLUCRAL BRACTS: small leaf-like structures comprising the involucre.

LINEAR DIVISIONS: bract divided into lobes that are cut more than half way to the base.

#214. Involucral bracts <colour>/
1. mostly green/
2. mostly red or purple pigmented/
3. outermost bracts green, sometimes with red pigment in centre; innermost bracts frequently purple or occasionally green/
4. outermost bracts green, with brown or red edges; innermost bracts almost transparent with red or brown central area/
5. with a green central portion and wide black margins/
6. with silvery grey central area and wide brown margins/

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families.

#215. Involucral bracts <shape>/
1. linear/
2. oblong/
3. lanceolate/
4. ovate/

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

LANCEOLATE:

OVATE: egg-shaped in outline and attached at the broad end.

#216. Involucral bracts <hairs>/
1. glabrous/
2. sparsely woolly/
3. densely woolly/

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families. Measured from base of involucre to tip of innermost bract.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

#217. Involucral bracts <whether hairs are glandular>/
1. with glandular hairs/
2. hairs non-glandular/

GLANDULAR: prosessing glands. These may be small sessile bulges that store or secrete fluids, or borne on the ends of glandular haris.

#218. Involucral bracts with number of rows/

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families, e.g. Cyperaceae.

Measured across the pressed involucre at the centre. INVOLUCRAL ROWS: these are one or more layers of bract leaves.

#219. Outer involucral bracts <height>/
mm high/

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families. Measured from base of involucre to tip of innermost bract.

#220. Outer involucral bracts <width>/
mm wide/

INVOLUCRE: one or more whorls of small leaves or bracts (phyllaries) standing close beneath a flower or flower structure. Common in the Daisy family, also occurs in other families, e.g. Cyperaceae.

Spikelets of grasses

#221. Spikelets <whether with pedicels>/
1. pedicellate/
2. sessile/

SPIKELETS: the basic unit of the inflorescence in the grasses and sedges, usually consisting of two glumes and one or more florets.

pedicelled: with pedicels, the stalks supporting the spikelets.

SESSILE: attached directly at the base, without a stalk.

#222. Spikelets <where disarticulating>/
1. disarticulating above the glumes/
2. disarticulating at the base of the spikelet/
3. disarticulating at the nodes of the main axis/

SPIKELET: the basic unit of the inflorescence in the grasses and sedges, usually consisting of two glumes and one or more florets.

DISARTICULATION: The point of disarticulation is where the spikelet breaks for seed dispersal.

GLUMES: the pair of bracts usually present at the base of the spikelet.

#223. Spikelets <compression>/
1. laterally compressed/
2. dorsoventrally compressed/

SPIKELET: the basic unit of the inflorescence in the grasses and sedges, usually consisting of two glumes and one or more florets.

This character records the plane of compression of a spikelet and is best determined in flowering material because the swelling of the fruit may alter the general shape. At anthesis, most spikelets are strongly compressed so that if one is placed on a flat surface it lies either on its side, the laterally compressed condition most common in Arctic taxa, or on its front or back, the dorsally compressed condition.

#224. Spikelets <shape in longitudinal section mainly for monocots>/
1. linear/
2. oblong/
3. lanceolate/
4. ovate/
5. oblanceolate/
6. obovate/

SPIKELET: the basic unit of the inflorescence in the grasses and sedges, usually consisting of two glumes and one or more florets.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

LANCEOLATE: lance or spear-shaped, much longer than wide, with the widest point below the middle.

OVATE: egg-shaped in outline and attached at the broad end.

OBLANCEOLATE: inversely lanceolate, with the attachment at the narrower end.

OBOVATE: inversely ovate, with the attachment at the narrow end.

#225. Spikelets <length>/
mm long/

SPIKELET: the basic unit of the inflorescence in the grasses and sedges, usually consisting of two glumes and one or more florets. This length does not include the length of any awns.

#226. Spikelets <width>/
mm wide/

SPIKELET: the basic unit of the inflorescence in the grasses and sedges, usually consisting of two glumes and one or more florets. The width is measured at the widest point.

#227. Florets per spikelet/

FLORET: the lemma and palea with its enclosed flower. The floret may be perfect (having female and male reproductive organs), pistillate (female only) or staminate (male only).

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret.

PALEA: the uppermost of the two bracts enclosing the flower in the grass floret.

Catkins

#228. Catkins flowering <relative time of catkin opening>/
1. before the opening of leaf buds/
2. with the opening of leaf buds/

CATKIN: an inflorescence consisting of a scaly-bracted spike of apetalous, unisexual flowers.

#229. <Catkins position on previous year’s shoot>/
1. catkins terminal on previous year’s shoot/
2. one to several catkins just below tip of previous year’s shoot/
3. numerous catkins just below tip of previous year’s shoot/

CATKIN: an inflorescence consisting of a scaly-bracted spike of apetalous, unisexual flowers

Male catkins

#230. Male catkins <density>/
1. densely flowered/
2. moderately densely flowered/
3. loosely flowered/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower. MALE CATKINS: those with the anthers.

RACHIS: axis of the catkin, bearing the flowers.

DENSELY FLOWERED: rachis not visible.

MODERATELY DENSELY FLOWERED: rachis partly visible.

LOOSELY FLOWERED: Rachis clearly visible.

#231. Male catkins <length>/
mm long/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers.

Length is the distance from the tip of the catkin to the lowermost flower.

#232. Male catkins <width of pressed catkins>/
mm wide/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers.

Width is measured at about the midpoint of the catkin.

#233. Male catkins <general shape>/
1. slender/
2. stout/
3. subglobose/
4. globose/
5. shape indeterminate <few flowered, shape not applicable>/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

SLENDER: greater than 3X longer than wide.

STOUT: less than 3X longer than wide.

SUBGLOBOSE: slightly longer than wide.

GLOBOSE: spherical in shape, like a ball.

#234. Male catkins peduncles <length>/
mm long/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

PEDUNCLE: the stalk, which can be naked or with 2–3 green bracts, is located between the lowermost flower on the catkin and the nearest (distal) leaf on the flowering branchlet.

NOTE: Peduncle length is measured from the lowermost flower on the catkin to the nearest (distal) leaf on the flowering branchlet or, if no flowering branchlet is present, to the stem.

#235. Male catkins <presence of leafy, flowering branchlets>/
1. borne on a flowering branchlet <terminating in a catkin>/
2. sessile/

FLOWERING BRANCHLET: a short, leafy shoot which terminates in a catkin. It bears three to several leaves that correspond to the leaves at the base of normal vegetative branchlets.

SESSILE: the catkin is attached directly to the branch.

#236. Male catkins flowering branchlets <length>/
mm long/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

FLOWERING BRANCHLET: a short, vegetative shoot on which the catkin is borne. It bears three to several leaves that correspond to the leaves at the base of normal vegetative branchlets.

NOTE: The flowering branchlet is measured from its point of attachment to the branch to the point of attachment of its uppermost (distal) leaf.

Female catkins

#237. Female catkins <density>/
1. densely flowered/
2. moderately densely flowered/
3. loosely flowered/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower. FEMALE CATKINS: structures bearing the carpels.

RACHIS: axis of the catkin, bearing the flowers.

DENSELY FLOWERED: rachis not visible.

MODERATELY DENSELY FLOWERED: rachis partly visible.

LOOSELY FLOWERED: Rachis clearly visible.

#238. Female catkins <length>/
mm long/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

Length is the distance from the tip of the catkin to the lowermost flower.

#239. Female catkins <width of pressed catkins>/
mm wide/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

Width is measured at about the midpoint of the catkin.

#240. Female catkins <general shape>/
1. slender/
2. stout/
3. subglobose/
4. globose/
5. shape indeterminate <few flowered, shape not applicable>/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

SLENDER: greater than 3X longer than wide.

STOUT: less than 3X longer than wide.

SUBGLOBOSE: slightly longer than wide.

GLOBOSE: spherical in shape, like a ball

#241. Female catkins peduncles <length>/
mm long/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

PEDUNCLE: the naked stalk (or with 2–3 green bracts) located between the proximal (lowermost) flower on the catkin and the first leaf on the flowering branchlet.

NOTE: Peduncle length is measured from the lowermost flower on the catkin to the nearest (distal) leaf on the flowering branchlet or, if no flowering branchlet is present, to the stem.

#242. Female catkins <presence of leafy, flowering branchlets>/
1. borne on a flowering branchlet/
2. sessile/

FLOWERING BRANCHLET: a short, vegetative shoot on which the catkin is borne at the end. It bears three to several leaves that correspond to the leaves at the base of normal vegetative branchlets.

SESSILE: the catkin is attached directly onto the stem.

#243. Female catkins flowering branchlets <length>/
mm long/

CATKIN: an inflorescence consisting of a dense spike or raceme of apetalous, unisexual flowers. It is measured from the tip of the inflorescence to the lowermost flower.

FLOWERING BRANCHLET: a short, vegetative shoot on which the catkin is borne. It bears three to several leaves that correspond to the leaves at the base of normal vegetative branchlets.

NOTE: The flowering branchlet is measured from its point of attachment to the branch to the point of attachment of its uppermost (distal) leaf.

Flowers

#244. Flowers per inflorescence <number>/

#245. Flowers <size>/
1. small, less than 5 mm in diameter or length/
2. medium-sized, 5–15 mm in diameter or length/
3. large, more than 15 mm in diameter or length/

#246. Flowers <whether stalked, borne on a pedicle that it a branch off the main flowering stem, the peduncle> /
1. sessile or subsessile/
2. borne on a stalk <pedicel>/

SESSILE: attached directly to the branch or stem, without a stalk.

#247. Flowers <sexuality>/
1. unisexual/
2. bisexual /

UNISEXUAL: a flower with either male or female reproductive organs, but not with both.

BISEXUAL: having both male and female reproductive organs on the same plant and in the same floral structure.

#248. Flowers <symmetry>/
1. actinomorphic /
2. zygomorphic/

SYMMETRY: the arrangement of the flower parts including sepals, petals, anthers and gynoecium.

ACTINOMORPHIC: regular, radially symmetrical; a flower in which all parts are similar in size and arrangement.

ZYGOMORPHIC: transversely symmetrical; floral parts with two symmetrical halves.

Male flowers of sedges

#249. Staminate flowers <visibility>/
1. inconspicuous <often concealed among pistillate flowers>/
2. conspicuous/

Female flower of sedges: floral scales

#250. Floral scales <pistillate scales in Carex relative length>/
1. shorter than the perigynium in fruit/
2. as long as the perigynium in fruit/
3. longer than the perigynium in fruit/

PERIGYNIUM (plural perigynia) the inflated sac surrounding the ovary in Carex or Kobresia that may be open or closed.

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#251. Floral scales <pistillate colour>/
1. brown/
2. black/
3. orange brown/
4. green/
5. tawny <dull yellowish-brown>/
6. light rose/
7. bicolour <darker toward the tip>/
8. pale grey <lead coloured>/
9. white or translucent/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#252. Floral scales <pistillate scale margins>/
1. with margins the same colour as the body of the scale/
2. with margins, and sometimes midvein paler in colour than the adjacent area of the scale/
3. with margins darker in colour than the midvein/
4. with margins paler than body of scale/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#253. Floral scales <pistillate orientation>/
1. reflexed/
2. not reflexed /

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

REFLEXED: bent back, and pointing downwards.

#254. Floral scales <pistillate shape at apex>/
1. truncate/
2. rounded/
3. retuse/
4. obtuse/
5. cuspidate/
6. acute/
7. lacerate/
8. deeply divided/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower. TRUNCATE: ending abruptly as though cut off transversely.

OVATE: rounded.

OBTUSE: blunt, at an angle more than 90 degrees but less than 180 degrees.

LANCEOLATE: shaped like the head of a spear, broadest near the base, narrowing towards the top.

CUSPIDATE: tipped with a sharp point.

ACUTE: tapering to a point; angle less than 45 degrees.

OBOVATE: inversely ovate, with the attachment at the narrow end.

ROUNDED: with a rounded apex.

RETUSE: with a shallow notch in a round or blunt apex.

LACERATE: cut or cleft irregularly, as if torn.

#255. Floral scales <shape>/
1. ovate/
2. lanceolate/
3. obovate/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower. TRUNCATE: ending abruptly as though cut off transversely.

#256. Floral scales <pistillate longevity>/
1. falling early <caducous>/
2. not falling early /

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#257. Floral scales <pistillate flowers>/
mm long/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#258. Floral scales <pistillate flowers>/
mm wide/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#259. Floral scales <hairiness on outer surface>/
1. glabrous/
2. hairy all over/
3. hairy mainly at apex/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

Floral bracts

#260. Floral bracts <colour> /
1. tawny <dull yellowish-brown>/
2. light rose/
3. brown/
4. black/
5. bicolour <darker toward tip>/
6. pale grey <lead coloured>/
7. orange brown/
8. green/
9. white or translucent/

FLORAL BRACT: a much reduced leaf subtending each flower.

GLABROUS: without hairs.

#261. Floral bracts apex/
1. entire/
2. lacerate/
3. deeply divided/
4. truncate/
5. rounded/
6. retuse/
7. obtuse/
8. cuspidate/
9. acute/

#262. Floral bracts <shape>/
1. ovate/
2. lanceolate/
3. obovate/

FLORAL BRACT: a much reduced leaf subtending each flower.

GLABROUS: without hairs.

ENTIRE: not toothed; margins smooth.

LACERATE: cut or cleft irregularly, as if torn. DEEPLY DIVIDED: cut more than 25%.

TRUNCATE: apical region flat.

#263. Floral bracts <shape, both male and female>/
1. widest at base/
2. widest at middle/
3. widest toward tip <apex>/

FLORAL BRACT: a much reduced leaf subtending each flower.

#264. Floral bracts <length>/
mm long/

#265. Floral bracts <width>/
mm wide/

#266. Floral bracts <hairiness on outer surface>/
1. glabrous/
2. hairy all over/
3. hairy mainly at apex/

#267. Floral bracts hairs <density>/
1. sparse/
2. moderately dense <surface 50% visible>/
3. very dense <surface completely obscured>/

FLORAL SCALE: any thin, scarious body, usually a degenerate leaf, sometimes of epidermal origin and subtending each flower.

#268. Floral bracts hairs <shape>/
1. straight/
2. wavy/

FLORAL BRACT: a much reduced leaf subtending each flower.

#269. Floral bracts <apex toothing>/
1. entire/
2. minute undulations/
3. divided into 3-lobes/

FLORAL BRACT: a much reduced leaf subtending each flower.

ENTIRE: not toothed; margins smooth.

DIVIDED INTO THREE LOBES: condition found in Betula.

Glumes of grasses

#270. <Glumes comment>/

GLUMES: the pair of bracts usually present at the base of the spikelet.

#271. First glume <length relative to the second glume>/
× the length of the second glume/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

SECOND GLUME: the upper such glume.

#272. First glume <length relative to the spikelet>/
× spikelet length/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

#273. First glume <length>/
mm long/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

#274. First glume <shape>/
1. linear/
2. oblong/
3. deltoid/
4. lanceolate/
5. ovate/
6. obovate/
7. oblanceolate/
8. transversely oblong/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel Sides.

DELTOID: triangular.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

OVATE: egg-shaped in outline and attached at the broad end.

OBOVATE: inversely ovate, with the attachment at the narrow end.

OBLANCEOLATE: inversely lanceolate, with the attachment at the narrower end.

#275. First glume <whether hairy>/
1. glabrous/
2. with trichomes/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

GLABROUS without hairs.

TRICHOMES: hairs or hair-like outgrowths, that may be scabrous (stiff) or pubescent (soft), on the surface of the glume.

#276. First glume margins <whether ciliate>/
1. glabrous/
2. ciliate/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

GLABROUS: without hairs.

CILIATE: fringed with hairs.

#277. First glume veins <number>/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet. Number of veins transversing the central point.

#278. First glume apex <shape of the apical region>/
1. caudate/
2. acuminate/
3. acute/
4. obtuse/
5. truncate/

FIRST GLUME: the lower of the pair of bracts usually present at the base of the spikelet.

CAUDATE: with a tail-like appendage.

ACUMINATE: apical region with sides somewhat concave, and which taper to an extended point.

ACUTE: apex formed by margins meeting at less than a 90 degree angle, sides more or less straight.

OBTUSE: rounded.

TRUNCATE: apical region flat.

#279. Second glume <length relative to the spikelet>/
1. 0.4 × as long as the spikelet or less/
2. 0.4–0.9 × as long as the spikelet/
3. as long, or longer than the spikelet/

SECOND GLUME: the upper of the pair of bracts usually present at the base of the spikelet.

#280. Second glume <length relative to the lowest floret, without lemma awn>/
1. shorter than the lowest floret/
2. almost as long as, or longer than, the lowest floret/

SECOND GLUME: the upper of the pair of bracts usually present at the base of the spikelet.

This measurement excludes the length of the lemma awn.

#281. Second glume <length>/
mm long/

SECOND GLUME: the upper of the pair of bracts usually present at the base of the spikelet.

#282. Second glume <shape>/
1. linear/
2. oblong/
3. deltoid/
4. lanceolate/
5. ovate/
6. elliptic/
7. oblanceolate/

SECOND GLUME: the upper of the pair of bracts usually present at the base of the spikelet.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

DELTOID: triangular.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

OVATE: egg-shaped in outline and attached at the broad end.

OBOVATE: inversely ovate, with the attachment at the narrow end.

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends.

OBLANCEOLATE: inversely lanceolate, with the attachment at the narrower end.

#283. Second glume <surface features>/
1. glabrous/
2. with trichomes/

SECOND GLUME: the upper of the pair of bracts usually present at the base of the spikelet.

GLABROUS: without hairs.

TRICHOMES: : hairs or hair-like outgrowths on the surface of the glume. They may be scabrous (stiff) or pubescent (soft).

#284. Second glume margins <whether ciliate>/
1. glabrous/
2. ciliate/

GLABROUS: without hairs.

CILIATE: fringed with hairs.

#285. Second glume veins <number>/

SECOND GLUME: the upper of the pair of bracts usually present at the base of the spikelet. Number of veins transferring the central point.

Rachilla of grasses

#286. Rachilla internode <length>/
mm long/

RACHILLA: the internal axis of the spikelet in grasses or a spike in sedges.

#287. Rachilla <width>/
mm wide/

RACHILLA: the internal axis of the spikelet in grasses and sedges.

#288. Rachilla internode <surface features>/
1. glabrous/
2. scabrous/
3. hairy/

RACHILLA: the internal axis of the spikelet in grasses or a spike in sedges.

GLABROUS: without hairs.

SCABROUS: rough to the touch because of small, bristly, prickle-hairs.

HAIRY: with long soft hairs.

#289. Rachilla <whether pronounced between the florets>/
1. pronounced between the florets/
2. not pronounced between the florets/

RACHILLA: the internal axis of the spikelet in grasses or a spike in sedges. If pronounced between the florets, the spikelet or spike may have a jointed, zig-zag appearance.

#290. Rachilla <whether terminating in a naked point or spikelet>/
1. terminating in a well-formed floret/
2. terminating in a vestigial floret/
3. extending beyond the uppermost floret <the terminal floret absent>/

RACHILLA: the internal axis of the spikelet in grasses or a spike in sedges.

VESTIGIAL: rudimentary and almost completely reduced; with only a vestige remaining.

Callus of grasses

#291. Callus <whether differentiated>/
1. differentiated/
2. not differentiated /

CALLUS: a hard thickening or protuberance; the thickened basal extension of the lemma in many grasses.

Caution. The callus may appear differentiated if hairs occur at the base of the lemma, but not on the callus.

#292. Callus <presence of hairs>/
1. glabrous/
2. hairy /

CALLUS: a hard thickening or protuberance; the thickened basal extension of the lemma in many grasses.

GLABROUS: without hairs.

#293. Callus hairs <length>/
mm long/

CALLUS: a hard thickening or protuberance; the thickened basal extension of the lemma in many grasses.

When the hairs are very curly, the lengths are only approximate.

#294. Callus hairs <length relative to the floret>/
1. shorter than the floret/
2. sub-equal to the floret/
3. longer than the floret/

CALLUS: a hard thickening or protuberance; the thickened basal extension of the lemma in many grasses. Subequal callus hairs occur in Calamagrostis canadensis.

Lemma of grasses

#295. Lemma <of fertile floret, length>/
mm long/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

#296. Lemma <of fertile floret, shape>/
1. linear/
2. oblong/
3. ovate/
4. lanceolate/
5. elliptic/
6. obovate/
7. oblanceolate/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBLONG: two to four times longer than wide with nearly parallel sides.

OVATE: egg-shaped in outline and attached at the broad end.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends.

OBOVATE: Inversely ovate, with the attachment at the narrow end.

OBLANCEOLATE: Inversely lanceolate, with the attachment at the narrower end.

#297. Lemma <of fertile floret, whether keeled>/
1. keeled/
2. rounded on the back/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

KEEL: a raised or pronounced ridge, often V-shaped.

#298. Lemma <whether inrolled>/
1. lemma not strongly incurved/
2. lemma strongly inrolled/

#299. Lemma surface <of fertile floret, luster>/
1. shiny/
2. dull/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

SHINY: similar to high gloss or semi-gloss paint.

DULL: surface reflection similar to that of flat paint.

#300. Lemma surface <of fertile floret, whether hairy>/
1. glabrous/
2. sparsely scabrous/
3. hairy/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

GLABROUS: without hairs.

SPARSELY scabrous: with a sparse covering of minute prickle-hairs.

HAIRY: with longer hairs on surface.

#301. Lemma surface with trichomes/
1. on veins only/
2. on and between the veins/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

TRICHOMES: hairs or hair-like outgrowths on the surface of the glume.

#302. Lemma veins <number>/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet. The number of veins transversing the central point.

Lemma apex and awn (grasses)

#303. Lemma apex <of fertile floret, shape of the apical region>/
1. caudate/
2. acuminate/
3. acute/
4. rounded/
5. truncate/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

CAUDATE: with a tail-like appendage.

ACUMINATE: apex with sides somewhat concave, and which taper to an extended point.

ACUTE: apex formed by margins meeting at less than a 90 degree angle, sides more or less straight.

ROUNDED: a rounded apex.

TRUNCATE: with a flat apex.

#304. Lemma apex <of fertile floret, apical modifications>/
1. entire/
2. erose/
3. lacerate/
4. bifid/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

Entire: not toothed, notched, or divided, as the continuous margins of some leaves.

EROSE: with the margin irregularly toothed, as if gnawed.

LACERATE: cut or cleft irregularly, as if torn.

BIFID: deeply two-cleft or two-lobed, usually from the tip.

#305. Lemma apex <of fertile floret, whether ciliate>/
1. not ciliate/
2. ciliate/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret. Data refer to the first lemma in the spikelet.

CILIATE: fringed with hairs.

#306. Lemma apex <of fertile floret, presence of apical appendages>/
1. awned/
2. awnless/

LEMMA: the lowermost of the two bracts enclosing the flower in the grass floret.

AWN: a substantial hair or bristle.

#307. Awn arising <of the fertile floret, position of the awn>/
1. from the tip/
2. from below the apex but above the middle/
3. from the middle or below <but not near the base>/
4. from just above the base/

AWN: a substantial hair or bristle.

#308. Awn <of fertile floret awn, length>/
mm long/

AWN: a substantial hair or bristle.

Palea of grasses

#309. Palea <of fertile floret, development>/
1. well developed/
2. vestigial/
3. absent/

PALEA: the uppermost of the two bracts enclosing the flower in the grass floret.

VESTIGIAL: rudimentary and almost completely reduced; with only a vestige remaining.

#310. Palea <length>/
mm long/

PALEA: the uppermost of the two bracts enclosing the flower in the grass floret.

#311. Palea <of fertile floret trichomes on the veins or keels>/
1. with glabrous veins/
2. with scabrous veins/
3. with hairy veins/

PALEA: the uppermost of the two bracts enclosing the flower in the grass floret.

GLABROUS: without hairs.

SCABROUS: rough to the touch because of small, bristly, prickle-hairs.

HAIRY: with longer, more slender hairs.

Flower

#312. Perianth <presence>/
1. present /
2. absent/
3. reduced to lodicules/
4. represented by bristles <both the calyx and the corolla>/

PERIANTH: the floral envelope that consist of calyx and corolla in most flowers but which is reduced to scale-like or bristle-like structures in some members the Cyperaceae and possibly lodicules in grasses.

LODICULE: Small scalelike processes usually two or three in number, at the base of the stamens in grass flowers.

#313. Perianth bristles <texture>/
1. silky white, or translucent/
2. dull white or yellowish/
3. inconspicuous/

#314. Epicalyx <presence of epicalyx>/
1. present/
2. absent /

EPICALYX:lobes of the hypathium, below the calyx that appear as a calyx tube; important in the Rosaceae.

#315. Epicalyx segments <length>/
mm long/

EPICALYX:lobes of the hypathium, below the calyx that appear as a calyx tube; important in the Rosaceae.

#316. Epicalyx segments <width>/
mm wide/

EPICALYX:lobes of the hypathium, below the calyx that appear as a calyx tube;

#317. Epicalyx segments <relative length>/
1. shorter than the calyx segments/
2. equal in length to the calyx segments/
3. longer than the calyx segments/

EPICALYX:lobes of the hypathium, below the calyx that appear as a calyx tube; important in the Rosaceae.

#318. Epicalyx segments <relative width>/
1. narrower than the calyx segments/
2. equal in width to a calyx segments/
3. wider than the calyx segments/

EPICALYX:lobes of the hypathium, below the calyx that appear as a calyx tube; important in the Rosaceae.

Calyx

#319. Calyx <presence>/
1. present /
2. absent/
3. modified to a pappus/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

SEPAL: an individual unit of the calyx; it is usually green and often hairy; in some plants the sepals are brightly coloured and assume the function of petals.

PAPPUS: peculiar modified calyx of a single floret, borne on the ovary (persisting in fruit), being plumose, bristlelike, and scales.

#320. Calyx base <shape at the point of attachment of the sepals to the pedicel for free sepals, particularly Cerastium species>/
1. narrowly angled <angle 20–45°>/
2. widely angled <angle 45–80°>/
3. rounded <the angle at the base 80–90°>/

#321. Calyx sepals <number of free or fused sepals with distinct tips>/

CALYX: the collective term for all the sepals; forms the outer perianth whorl. SEPALS: an individual unit of the calyx; it is usually green and often hairy; in some plants the sepals are brightly coloured and assume the function of petals.

#322. Calyx sepals <fusion>/
1. free/
2. fused/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

FREE: sepals independent of one another.

FUSED: sepals joined to one another, usually forming a tube (calyx).

#323. Calyx sepals <length in mm> <measure the length of a sepal: don't measure if calyx is a pappus>/
mm long/

#324. Calyx <sepals or calyx segments width>/
mm wide/

Character developed for Cerastium.

#325. Calyx <sepals, or episepals> <colour>/
1. green/
2. yellow/
3. brown/
4. purple/
5. red/
6. blue/
7. black/
8. white or translucent/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

#326. Calyx <or sepals> <texture>/
1. herbaceous /
2. scarious/
3. petaloid/
4. fleshy/
5. leathery/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

SCARIOUS: a thin, dry, non-green membranous structure.

#327. Calyx <or calyx> <accrescent>/
1. accrescent/
2. non-accrescent /

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

ACCRESCENT: expanding and growing larger after flowering; bigger in fruit, than in flower, eg. bladder campion.

#328. Calyx <or sepals> <glands>/
1. with sessile glands/
2. without sessile glands/

#329. Calyx <or sepals> <hairs>/
1. glabrous/
2. hairy/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

GLABROUS: without hairs.

#330. Calyx <surface pubescence>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. woolly/
7. hairs short-silky <less than 0.5 mm>/
8. hairs long-silky <greater than 0.5 mm>/

SEPALS: an individual unit of the calyx; it is usually green and often hairy; in some plants the sepals are brightly coloured and assume the function of petals.

PUBERULENT: minutely short hairy or downy.

PUBESCENT: covered with short, fine hairs, downy.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: bearing short, dense, soft, wooly hairs, that are matted or tangled.

WOOLLY: bearing long, dense, soft hairs, that are tangled.

SILKY: straight, appressed, shiny hairs.

#331. Calyx <or sepal> hairs <whether glandular>/
1. glandular/
2. non-glandular/

GLANDULAR HAIRS: hairs with a swollen cell that may secrete, often sticky, material.

#332. Calyx hairs <colour>/
1. white or translucent/
2. brown/
3. black/
4. transparent with red cross-walls/

#333. Calyx <sepals, presence of glands>/
1. with stalked glands/
2. without stalked glands /

GLAND: a swollen cell or group of cells that may secrete often sticky material.

#334. Calyx <sepal margins, whether ciliate>/
1. margins ciliate/
2. margins without cilia /

#335. Calyx <shape when sepals are fused>/
1. tubular/
2. bell-shaped/
3. bilabiate/
4. funnel-form/
5. rotate/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

TUBULAR: shaped like a cylindrical tube.

BELL-SHAPED: a tube that is narrow at one end and flared out towards the other end.

BILABIATE: with two lips; each lip may be lobed or toothed.

FUNNEL-FORM: gradually widening from base to apex; funnel-shaped.

ROTATE: disc-shaped; flat and circular, with widely spreading lobes and little or no tube.

#336. Calyx <margin, when sepals fused>/
1. unlobed/
2. 2-lobed/
3. 3-lobed/
4. 4-lobed/
5. 5-lobed/

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

NOTE: If there only appears to be one perianth whorl in a flower it is usually considered to be the calyx.

LOBED: bearing lobes which are cut less than half way to the base or midvein.

#337. <Pappus>/
1. with a single row of hairs/
2. with a double row of hairs/

PAPPUS: peculiar modified calyx of a single floret, borne on the ovary (persisting in fruit), being plumose, bristlelike, and may have scales.

DOUBLE PAPPUS: two distinct lengths of pappus filaments.

#338. Ray floret pappus <length>/
mm long/

RAY FLORET: ligulate floret on Asteraceae head.

PAPPUS: peculiar modified calyx of a single floret, borne on the ovary (persisting in fruit), being plumose, bristlelike, and may have scales.

#339. Disk floret pappus <length>/
mm long/

DISK FLORET: tubular floret on Asteraceae head.

PAPPUS: peculiar modified calyx of a single floret, borne on the ovary (persisting in fruit), being plumose, bristlelike, and may have scales.

#340. Calyx teeth <whether equal in a fused calyx>/
1. equal or nearly so/
2. sub-equal or unequal/

CALYX TEETH: shape pointed lobes at the top of the calyx sepals.

#341. Calyx teeth <length>/
mm/

CALYX TEETH: shape pointed lobes at the top of the calyx sepals.

#342. Calyx teeth <whether glandular verrucose, in a fused calyx>/
1. without or with few glandular verrucae/
2. with abundant glandular verrucae/

VERRUCOSE: covered with verrucae.

VERRUCAE: wart-like outgrowths.

Petals

#343. Petals <presence>/
1. present /
2. absent/

PETAL: an individual segment of the corolla.

#344. Petals <free, number>/

PETAL: an individual segment of the corolla.

#345. Petals <fusion>/
1. free <to the base>/
2. fused <all petals, or all but one position in the corolla (ray florets) joined at least at base>/
3. both free and fused <as in pea flowers>/

PETAL: an individual segment of the corolla.

FREE: not attached to other petals at the base.

FUSED: petals are joined at least at the base, but can be fused to the tip of the petal.

#346. Petals <colour>/
1. green/
2. white/
3. yellow/
4. orange/
5. red/
6. pink/
7. purple/
8. blue/
9. brown/

PETAL: an individual segment of the corolla.

COROLLA: the collective name for all the petals; the inner perianth whorl.

#347. Petals <markings>/
1. with contrasting markings /
2. without contrasting markings /

CONTRASTING MARKINGS: distinctly different lines or patches of colour.

Free petals

#348. Petals <free> <shape>/
1. elliptic/
2. ovate/
3. obovate/
4. lanceolate/
5. oblanceolate/
6. spatulate/
7. linear/
8. obtriangular/

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends.

OVATE: egg-shaped in outline and attached at the broad end.

OBOVATE: inversely ovate, with the attachment at the narrow end.

LANCEOLATE: lance-shaped; much longer than wide, with the widest point below the middle.

OBLANCEOLATE: inversely lanceolate, with the widest point above the middle.

SPATULATE: spoon-shaped, with a rounded blade above gradually tapering to the base.

LINEAR: resembling a line; long and narrow with more or less parallel sides.

OBTRIANGULAR: shaped like an inverted triangle.

#349. Petals <free> <apex>/
1. unlobed/
2. 2-lobed/
3. 3-lobed/
4. 4-lobed/
5. 5-lobed/

COROLLA: the collective name for all the petals; the inner perianth whorl.

LOBED: bearing lobes which are cut less than half way to the base or midvein.

#350. Petals <free> <apex>/
1. tipped with an awn-like bristle/
2. not tipped with an awn-like bristle /

#351. Petals <length in mm>/
mm long/

PETAL: an individual segment of the corolla.

COROLLA: the collective name for all the petals; the inner perianth whorl.

#352. Petals <width>/
mm wide/

#353. Petals <spurred>/
1. spurred/
2. not spurred /

PETAL: an individual segment of the corolla.

COROLLA: the collective name for all the petals; the inner perianth whorl.

SPURRED: bearing a spur or spurs (a hollow, slender, sac like appendage of a petal).

#354. Petals <length relative to sepals>/
1. shorter than the calyx/
2. same length as the calyx/
3. longer than the calyx /

CALYX: the collective term for all the sepals; forms the outer perianth whorl.

Fused petals

#355. Corolla <fused, number of lobes>/
1. unlobed/
2. 2-lobed/
3. 3-lobed/
4. 4-lobed/
5. 5-lobed/

COROLLA: the collective name for all the petals; the inner perianth whorl.

#356. Corolla <shape>/
1. tubular/
2. rotate/
3. cup-like/
4. bilabiate/
5. campanulate <bell-shaped>/
6. urceolate/
7. funnel-form/
8. papilionaceous/
9. ligulate/

This character only applies to petals that are fused.

COROLLA: the collective name for all the petals; the inner perianth whorl.

TUBULAR: with the form of a tube or cylinder.

ROTATE: disc-shaped; flat and circular, with widely spreading lobes and little or no tube.

BILABIATE: two-lipped; each lip may be lobed or toothed.

CAMPANULATE: bell-shaped.

URCEOLATE: pitcher or vase-like; hollow and contracted near the mouth like a pitcher or urn.

BELL-SHAPED: a narrow tube and spreading lobes.

FUNNEL-FORM: gradually widening from base to apex; funnel-shaped

PAPILIONACEOUS: consisting of 5 petals with a banner petal (the uppermost one, with a fold along the middle), two wing petals, and two fused petals that form the keel.

LIGULATE: corolla of the ray florets tubular at the base, prolonged on the outer side into a flat, more or less strap-shaped organ that may appear as a single petal.

#357. Corolla keel <whether tipped>/
1. blunt/
2. with a pointed tip/

COROLLA: collective name for all the petals; the inner perianth whorl.

KEEL: part of the legume flower formed by and two fused petals.

#358. Wing auricles <free or united>/
1. free from each other, blunt, shorter than the claw/
2. united, linear, nearly equal to the claw/

WINGS: Two outermost petals of the papilionaceous flower.

AURICLES: Flaps at the base of the wing petals.

CLAW: Narrow basal part of wing petal.

BANNER: upper petal of papilionaceous flower with a fold in the middle.

KEEL: two lower petals join to form the keel.

#359. Anthers <whether present>/
1. present and probably functional <implicit>/
2. absent, or not functional/

Ray florets

#360. Ray florets <number of>/

RAY FLORET:outer modified floret of some Asteraceae with an expanded or strap-like part to the corolla.

LIMB: the expanding flat part of the corolla.

#361. Ray florets limb <length>/
mm long/

#362. Ray florets limb <width>/
mm wide/

Stamens

#363. Stamens <number>/

STAMENS: the male reproductive organ of a flower, consisting of an anther (pollen bearing portion) and filament (stalk that supports the anther).

#364. Stamens <relative lengths>/
1. filaments markedly unequal in length/
2. filaments all equal in length /

STAMEN: the male reproductive organ of a flower, consisting of an anther and filament.

#365. Stamens <attachment to corolla>/
1. fused to the corolla/
2. free of the corolla /

STAMEN: the male reproductive organ of a flower, consisting of an anther and filament.

NOTE: In some flowers, the stamens are fused to the corolla (petals), while in others they are completely free from the petals.

#366. Stamens filaments <hairiness>/
1. glabrous/
2. hairy all over/
3. hairy on lower half/
4. hairy at base only/

STAMENS: the male reproductive organ of a flower, consisting of an anther (pollen bearing portion) and filament (stalk that supports the anther).

GLABROUS: without hairs.

#367. Anthers <colour>/
1. purple <dark purple>/
2. purple becoming yellow/
3. reddish, <or brownish> becoming yellow/
4. yellow/
5. blue/

ANTHER: is the pollen bearing structure at the tip of the filament.

#368. Anthers <shape, when dehiscent>/
1. ellipsoid/
2. stout-cylindrical <less than 3x as long as wide>/
3. slender-cylindrical <greater than 3x as long as wide>/
4. ovoid/
5. subglobose/
6. triangular/

ANTHER: is the pollen bearing portion of the male reproductive organ of a flower.

DEHISCENT: opening at maturity to release the pollen.

ELLIPSOID: football-shaped; a three dimensional structure that is elliptic in long section and circular in cross section.

CYLINDRICAL: cylinder-shaped, elongate and round in cross-section.

OVOID: egg-shaped.

SUBGLOBOSE: almost spherical in shape, like a ball.

TRIANGULAR:shaped like an equilateral or Isosceles triangle.

#369. Anthers <axis shape after dehiscence>/
1. axis straight/
2. axis twisted/

ANTHER: is the pollen bearing portion of the male reproductive organ of a flower.

DEHISCENT: opening at maturity to release the pollen.

AXIS: the central line-like tissue of the anther.

#370. Anthers <including any apiculum (apical) and basal appendages length of dry, empty anthers>/
mm long/

ANTHER: is the pollen bearing portion of the male reproductive organ of a flower. APICULUM: terminated by a short, sharp point.

Nectaries

#371. Nectaries <in bisexual flowers>/
1. present/
2. absent /

NECTARY: a nectar secreting gland, often appearing as a protruberance, scale, or pit.

#372. Male flowers abaxial <between stamens and bract> nectaries <number>/
1. absent/
2. one/
3. two/

ABAXIAL: the side of a lateral organ away from the axis.

NECTARY: a nectar secreting gland, often appearing as a protruberance, scale, or pit.

#373. Male flowers adaxial <between stamens and rachis> nectaries <number>/
1. one/
2. two/
3. several/

ADAXIAL: the side of a lateral organ adjacent to the axis.

NECTARY: a nectar secreting gland, often appearing as a protruberance, scale, or pit.

#374. Male flowers adaxial <between stamens and rachis> nectaries <shape>/
1. slender-rod/
2. broad-rod/
3. square/
4. ovate/
5. half-cup-shaped/

ADAXIAL: the side of a lateral organ adjacent to the axis.

NECTARY: a nectar secreting gland, often appearing as a protruberance, scale, or pit.

SLENDER-ROD: length of nectary 4 or more times width.

BROAD-ROD: length of nectary 4 or more times width.

SQUARE: about as long as wide.

OVATE: egg-shaped, broadest at the base, narrowed towards the tip.

HALF-CUP SHAPED: base of nectary curved around the stamens.

#375. Male flowers adaxial <between stamens and rachis> nectaries <length>/
mm long/

ADAXIAL: the side of a lateral organ adjacent to the axis.

NECTARY: a nectar secreting gland, often appearing as a protruberance, scale, or pit.

#376. Male flowers adaxial <between stamens and rachis> nectaries <coalescence>/
1. separate/
2. partially connected at base/
3. cup-shaped/

ADAXIAL: the side of a lateral organ adjacent to the axis.

NECTARY: a nectar secreting gland, often appearing as a protruberance, scale, or pit.

Female flowers of Salix

#377. Female flowers adaxial <between stamens and rachis> nectaries <presence and number>/
1. absent/
2. one/
3. two/

FLORAL BRACT: a reduced leaf or scale-like structure at the base of a flower.

NECTARY: the organ which produces the nectar.

OVARY: the portion of the pistil that contains the ovules (immature seeds).

PISTIL: the female reproductive organ, comprised of ovary, style and stigma.

RACHIS: axis of the catkin, bearing the flowers.

STYLE: the usually elongated portion of the pistil between the stigma and the ovary. STIGMA: the part of the pistil which receives the pollen.

STIPE: the stalk of the pistil.

#378. Female flowers adaxial <between stamens and rachis> nectaries <lobeing>/
1. unlobed/
2. two or more lobes/

FLORAL BRACT: a reduced leaf or scale-like structure at the base of a flower.

NECTARY: the organ which produces the nectar.

OVARY: the portion of the pistil that contains the ovules (immature seeds).

PISTIL: the female reproductive organ, comprised of ovary, style and stigma. STIGMA: the part of the pistil which receives the pollen.

STIPE: the stalk of the pistil.

RACHIS: axis of the catkin, bearing the flowers.

STYLE: the usually elongated portion of the pistil between the stigma and the ovary.

#379. Female flowers adaxial <between stamens and rachis> nectaries <between ovary and rachis> <shape>/
1. slender-rod/
2. broad-rod/
3. square/
4. ovate/

ADAXIAL: the side facing the main axis of the plant.

NECTARY: the organ which produces the nectar.

OVARY: the portion of the pistil that contains the ovules (immature seeds).

PISTIL: the female reproductive organ, comprised of ovary, style and stigma.

RACHIS: axis of the catkin, bearing the flowers.

SLENDER-ROD: length 4 or more times width.

BROAD-ROD: length 2–3 times width.

SQUARE: approximately as long as wide.

OVATE: egg-shaped in outline and attached at the broad end.

#380. Female flowers adaxial <between stamens and rachis> nectaries <length>/
mm long/

ADAXIAL: the side facing the main axis of the plant.

NECTARY: the organ which produces the nectar.

OVARY: the portion of the pistil that contains the ovules (immature seeds).

#381. Female flowers adaxial <between stamens and rachis> nectaries <relative length>/
1. shorter than stipes/
2. equal to stipes/
3. longer than stipes/

ADAXIAL: the side facing the main axis of the plant.

NECTARY: the organ which produces the nectar.

OVARY: the portion of the pistil that contains the ovules (immature seeds).

#382. Female flowers nectaries <coalescence> /
1. separate/
2. partially connected/

COALESCENCE: fusion between like parts.

FLORAL BRACT: a reduced leaf or scale-like structure at the base of a flower.

NECTARY: the organ which produces the nectar.

OVARY: the portion of the pistil that contains the ovules (immature seeds).

PISTIL: the female reproductive organ, comprised of ovary, style and stigma.

RACHIS: axis of the catkin, bearing the flowers.

STYLE: the usually elongated portion of the pistil between the stigma and the ovary. STIGMA: the part of the pistil which receives the pollen.

STIPE: the stalk of the pistil.

Receptacles

#383. Receptacle <height>/
mm high/

RECEPTACLE: the portion of the stem upon which the flower or inflorescence is borne.

Carpels and ovaries

#384. Carpels <fusion>/
1. monomerous/
2. apocarpous/
3. partly fused/
4. syncarpous <completely fused>/

CARPEL: female reproductive organ, consisting of a stigma, style, and ovary.

MONOMEROUS: having a single carpel.

APOCARPOUS: having two or more carpels that are free from one another.

SYNCARPOUS: having two or more carpels that are fused.

#385. Carpels <number>/

CARPEL: female reproductive organ, consisting of a stigma, style, and ovary.

NOTE: In order to determine the number of carpels it may be necessary to take a cross section of the flower and count the number of segments. Each segment usually is one carpel.

#386. Gynoecia <position, when carpels fused or appearing monomerous>/
1. superior/
2. partly inferior/
3. inferior/

GYNOECIA: the female organs of a flower.

SUPERIOR (hypogynous): having the flower parts attached below the base of the ovary and free from it.

PARTLY INFERIOR (perigynous): having the flower parts attached to the edge of a cup shaped receptacle.

INFERIOR (epigynous): having the flower parts attached at or above the top of the ovary.

#387. Stipes <length>/
mm long/

STIPE: the stalk of an ovary.

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

#388. Ovaries <shape of superior ovaries>/
1. inverse club-shaped <obclavate>/
2. pear-shaped <pyriform>/
3. ovate <egg-shaped>/
4. inverse turnip-shaped/
5. urceolate/
6. oblong/
7. elliptic/
8. subglobose/
9. clavate/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

OBCLAVATE: inverse club-shaped.

URCEOLATE: hollow and narrowed at the mouth like an urn or pitcher.

OBLONG: rectangular in outline.

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends. CLAVATE: club-shaped.

#389. Ovary <beak shape>/
1. gradually tapering to style <no evident transition>/
2. slightly bulged at the base of the style <slight curve to style>/
3. abruptly tapering to style/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

BEAK: narrowed unswollen portion of the ovary.

STYLE is the slender stalk that connects the stigma to the ovary.

STIGMA: the part of the pistil which receives the pollen.

#390. Ovaries <general hairiness>/
1. glabrous/
2. hairy/

OVARY: the portion of the pistil that contains the ovules (immature seeds). GLABROUS: without hairs.

#391. Ovary <pubescence type>/
1. puberulent/
2. pubescent/
3. pilose/
4. villous/
5. tomentose/
6. hairs short-silky <less than 0.5 mm>/
7. hairs long-silky <greater than 0.5 mm>/
8. strigose/

OVARY: the swollen portion of the pistil that contains the ovules (immature seeds).

PUBERULENT: minutely pubescent or downy.

PUBESCENT: covered with short, fine hairs, downy.

PILOSE: bearing sparse, long, soft, straight, shaggy hairs.

VILLOUS: bearing dense or moderately dense, long, soft, shaggy, but not matted hairs.

TOMENTOSE: bearing short, dense, soft, wooly hairs, that are matted or tangled.

SILKY: straight, appressed, shiny hairs.

STRIGOSE: covered with straight, stiff hairs or bristles.

#392. Ovary hair <density>/
1. in patches or streaks/
2. sparse/
3. moderately dense/
4. dense/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

SPARSE: surface mostly visible.

MODERATELY DENSE: surface about 50% visible.

DENSE: surface completely obscured.

#393. Ovary hair <colour>/
1. white, or translucent/
2. a mixture of white, or translucent, and rust-coloured/
3. brown/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

#394. Ovary hair <orientation to surface>/
1. appressed/
2. spreading/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

APPRESSED: hair pressed close or flat against the branchlet.

SPREADING: hair standing out from the surface.

#395. Ovary hair <shape>/
1. straight/
2. wavy/
3. crinkled/
4. branched/
5. stellate/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

WAVY: in loosely spaced waves.

CRINKLED: in tightly spaced waves.

BRANCHED: hairs dividing irregularly.

STELLATE: hairs radiating like the points of a star.

#396. Ovary hair <type>/
1. flattened/
2. ribbon-like/
3. cylindrical/
4. with thickened bases/

OVARY: the expanded basal portion of the pistil that contains the ovules (immature seeds).

FLATTENED: hair edges appear thickened, and hairs sometimes refract light.

RIBBON-LIKE: hair edges flat and refracting light.

Styles and stigmas

#397. Styles <presence>/
1. present /
2. absent/

STYLE: the narrowed portion of the gynoecium connecting the stigma (top portion) to the ovary.

#398. Styles <fusion, when styles greater than one per ovary>/
1. free/
2. partially fused/
3. completely fused/

Character significant in Potentilla.

STYLE: the narrowed portion of the pistil (gynoecium) connecting the stigma (top portion) to the ovary.

FREE: not attached to other styles.

PARTIALLY FUSED: one style appears to emerge from the ovary and then branched into two or more.

#399. Styles <basal portion> /
1. base an enlarged bulge, separated from the top of the achene <Eleocharis>/
2. base not enlarged, continuous with the achene /
3. base enlarged and attached to the achene /

STYLE: the narrowed portion of the pistil (gynoecium) between the stigma to the ovary.

#400. Styles <relative length, used in Cyperaceae>/
1. thick and short/
2. slender, not extending beyond the beak/
3. slender, extending beyond the beak/
4. long and thick/

#401. Styles <number, when carpels fused>/

STYLE: the narrowed portion of the gynoecium connecting the stigma (top portion) to the ovary.

#402. Styles <length>/
mm long/

STYLE is the narrowed portion of the pistil (gynoecium) connecting the stigma (top portion) to the ovary.

#403. Style <shape or angle of the sides of the style, significant in Potentilla>/
1. conical <tapering sides>/
2. straight <sides parallel>/

#404. Style basal portion <surface>/
1. smooth <without any roughness>/
2. covered with short papillae, less than 0.1 mm high/
3. covered with long papillae, 0.1 mm high or higher/

#405. Stigmas per style <number>/

STIGMA: the top portion of the gynoecium which is receptive to pollen.

#406. Stigmas <shape>/
1. slender-cylindrical <e.g. Salix alaxensis>/
2. broad-cylindrical <e.g. Salix reticulata>/
3. two plump lobes with continuous stigmatic surface <connecting all 4 lobes, e.g. Salix herbacea>/
4. modified into a receptive area near the pollinia and the top of the column <Orchidaceae>/

STIGMA: the top portion of the pistil which is receptive to pollen.

CYLINDRICAL: cylinder shaped, elongate and round in cross-section.

#407. Stigmas lobes <length>/
mm long/

STIGMA: the top portion of the pistil which is receptive to pollen.

#408. Stigmas <shape in apocarpous carpels>/
1. plate shaped/
2. capitate <rounded and head-like>/
3. receptive surface at the end of an otherwise unmodified style/

Placentation

#409. Placentation <when carpels fused>/
1. axile /
2. parietal/
3. apical/
4. basal/
5. free central/

PLACENTATION: the arrangement or configuration of the placentae (the position(s) on the ovary bearing ovules).

OVARY: the expanded basal portion of the pistil that contains the ovules.

OVULE: an immature seed.

LOCULE: a cavity containing the seed(s)in an ovary.

AXILE: ovules attached to the central axis of an ovary with two or more locules.

PARIETAL: ovules positioned along the walls of the ovary.

APICAL: ovules located at the tip of the ovary.

BASAL: ovules located at the base of the ovary.

FREE CENTRAL: ovules attached to a central column which is free from the ovule wall except at the base.

Ovules

#410. Ovules <number per gynoecium>/

OVULE: an immature seed.

CARPEL: one of the leaf-like units that make up a compound pistil.

CAPSULE: a dry, dehiscent fruit composed of more than one carpel.

Perigynia

#411. Fruit <whether surrounded>/
1. surrounded by a perigynium/
2. not surrounded by a perigynium /

PERIGYNIUM (plural perigynia) the inflated sac surrounding the ovary in Carex or Kobresia that may be open or closed.

#412. Perigynia <whether fused>/
1. open on one side/
2. with a slit running down the beak on the abaxial side through which the style protrudes/
3. fused to the apex except for a small aperture through which the style protrudes/

#413. Perigynia <shape>/
1. globose, or subglobose/
2. lanceolate/
3. broadly ovate/
4. oblong/
5. obovate/
6. elliptic/

PERIGYNIUM (plural perigynia) the inflated sac surrounding the ovary in Carex or Kobresia that may be open or closed.

GLOBOSE: rounded like a ball.

SUB-LANCEOLATE longer than wide, relatively narrow and tapering to a point at the top.

BROADLY-OVATE: longer than wide, distinctly rounded with the widest point below the middle.

OBLONG: rectangular in outline.

OBOVATE: inversely ovate, with the attachment at the narrow end.

ELLIPTIC: in the shape of an ellipse, or a narrow oval; broadest at the middle and narrower at the two equal ends.

#414. Perigynia <height>/
mm long/

HEIGHT: measured from the base of the perigynia excluding the length of any stipe.

#415. Perigynia <width>/
mm wide/

#416. Perigynia <with or without a stalk, stipe>/
1. contracted at the base into a stipe-like structure/
2. sessile/

SESSILE: without a stalk.

#417. Perigynia <position of at least the lower perigynia relative to stem of spike>/
1. erect or ascending/
2. reflexed/
3. spreading at maturity/

ERECT: vertical or almost vertical.

ASCENDING: within the range of angles between erect and divergent.

REFLEXED: bent back, and pointing downwards.

#418. Perigynia <colour>/
1. black/
2. straw-coloured/
3. golden brown/
4. brown/
5. green/
6. whitish/

#419. Perigynia <texture>/
1. membranous <thin and almost translucent>/
2. leathery/

MEMBRANOUS: thin, soft flexible, and more or less translucent, like a membrane.

LEATHERY: leather-like.

#420. Perigynia <surface luster>/
1. surface glossy/
2. surface dull/

glossy: with a shine similar to full gloss paint.

DULL lacking a shine, similar to flat paint.

#421. Perigynia <indumentum>/
1. glabrous/
2. hairy/
3. scabrous/

#422. Perigynia <whether tuberculate>/
1. tuberculate/
2. glandular/
3. papillose/

TUBERCULATE: with knobby projections.

GLANDULAR: with small bulges that store or secrete fluids.

PAPILLOSE: with minute, nipple-shaped structures.

#423. Perigynia <visibility of nerves on the abaxial, ventral surface>/
1. strongly nerved/
2. faintly nerved/
3. appearing nerveless/

NERVE: vein and associated tissue.

STRONGLY NERVED: with nerves clearly visible.

#424. Perigynia <whether inflated>/
1. inflated/
2. not inflated /

INFLATED: puffed out, loosely fitting around the achene.

#425. Perigynia <whether keeled>/
1. not keeled /
2. with 2 keels/
3. with 3 keels/

KEEL: a raised or pronounced ridge that is often V-shaped.

#426. Perigynia apices/
1. beaked with a long beak/
2. beaked with a short beak/
3. merely conical or rounded <almost beakless>/
4. without a beak/

BEAK: a firm prolonged slender tip.

#427. Perigynia apex <beak>/
1. oblique, becoming slightly bidentate/
2. deeply bidentate/
3. not bidentate /

OBLIQUE: slanting.

BIDENTATE: having two teeth.

Fruit

#428. Fruit <calyx persisting>/
1. with calyx persisting/
2. without calyx persisting /
3. surrounded by perianth persisting as bristles/

This character refers to whether the sepals are present or not when the plant is in the fruiting stage.

CALYX: the collective term for all the sepals; the outer perianth whorl.

PAPPUS: peculiar modified calyx of a single floret, borne on the ovary (persisting in fruit), being plumose, bristlelike, and scales.

#429. Fruit <type>/
1. a follicle/
2. a legume/
3. a loment/
4. a capsule/
5. a silique/
6. an achene/
7. a nut/
8. a berry/
9. a drupe/
10. an aggregate of drupelets/
11. a schizocarp/
12. a caryopsis/
13. a samara/
14. an aggregate of nutlets/
15. an aggregate of follicles/
16. a mericarp/

FRUIT: a ripened ovary and any other structures which are attached and ripen with it.

FOLLICLE: a dry, dehiscent (opening at maturity or when ripe) fruit, derived from one carpel, that splits along a single side.

LEGUME: a dry, dehiscent fruit derived from a one carpel and usually opening along two lines of dehiscence, as does a pea pod.

LOMENT: a pod constricted between the seeds, and breaking into 1-seeded portions.

CAPSULE: a dry, dehiscent fruit derived from more than one carpel and splitting into as many segments as there are carpels.

SILIQUE: a dry, dehiscent fruit, with two carpels separated by a septum as in the Draba family. Fruits split from the bottom upwards.

ACHENE: a small, dry, indehiscent one seeded fruit with tight thin pericarp.

NUT: a hard, dry, indehiscent fruit, the product of more than one carpel often with a single seed.

BERRY: a fleshy fruit, with several or many seeds, as a tomato.

DRUPE: a fleshy, indehiscent fruit with a stony endocarp surrounding the single seed, such as a peach.

AGGREGATE OF FLESHY DRUPELETS: very small drupes that maybe aggregated into a fleshy fruit such as a raspberry

SCHIZOCARP: a fruit that splits up without the opening of the carpel into one-seeded units (mericarps).

CARYOPSIS: the indehiscent fruit in grasses.

SAMARA: a winged 1-seeded indehiscent achene or nut.

NUTLET: one-seeded, dry, indehiscent fruit, the product of an apocarpous carpel.

MERICARP: a fruit composed of more than one schizocarp.

#430. Fruit <shape>/
1. spherical/
2. ellipsoid/
3. ovoid/
4. elongate-cylindrical/
5. obovate/
6. oblong/
7. clavate/
8. urceolate/
9. conical/

FRUIT: a ripened ovary and any other structures which are attached and ripen with it.

SPHERICAL: globe-shaped.

ELLIPSOID: In the shape of a narrow oval; broadest in the middle, and narrow at the two ends.

OVOID: egg-shaped.

ELONGATE-CYLINDRICAL: cylinder-shaped; elongate and round in cross section.

DISTINCTLY FLAT: compressed.

OBOVATE: inversely ovate, with the attachment at the narrow end.

OBLONG: rectangular in outline.

CLAVATE: club-shaped.

U: urn-shaped. rounded and wider at the base, tapering towards the apex.

#431. Fruit <whether flattened>/
1. not distinctly flattened/
2. distinctly flattened/

#432. Fruit <dehiscence>/
1. dehiscent/
2. indehiscent/
3. schizocarpous/

FRUIT: a ripened ovary and any other structures that are attached and ripen with it.

DEHISCENT: opening at maturity or when ripe.

INDEHISCENT: not opening at maturity along definite lines or by pores.

SCHIZOCARPOUS: opening at maturity into separate one-seeded segments (carpels) at maturity, instead of along dehiscent lines, eg. maple seeds, umbels.

#433. Fruit <how the fruit opens>/
1. opening with apical pores/
2. opening with teeth at the top of the capsule/
3. splitting to the base into separate segments/

#434. Fruit <length or height>/
mm long/

FRUIT: a ripened ovary and any other structures that are attached and ripen with it. A fruit has two scars: one the point of attachment to the plant, the other the point of attachment of the style.

LENGTH OR HEIGHT: the distance between the point of attachment to the plant and the point of attachment of the style.

#435. Fruit <width>/
mm wide/

FRUIT: a ripened ovary and any other structures that are attached and ripen with it. A fruit has two scars: one the point of attachment to the plant, the other the point of attachment of the style.

WIDTH: the greatest dimension at right angles to the line between the point of attachment to the plant and the point of attachment of the style.

#436. Fruit <whether stalked>/
1. sessile/
2. stalked/

#437. Fruit stalk <peduncle length in fruit>/
mm long/

#438. Fruit <texture at maturity>/
1. fleshy <at least partly>/
2. dry/

FRUIT: a ripened ovary and any other structures that are attached and ripen with it.

FLESHY: thick and pulpy; succulent.

DRY: without water.

#439. Fruit <colour>/
1. yellowish/
2. black/
3. brown/
4. red/
5. purple/
6. blue/
7. green at maturity/
8. golden brown/
9. straw coloured/

FRUIT: a ripened ovary and any other structures that are attached and ripen with it.

#440. Fruit <hairs>/
1. glabrous /
2. hairy/
3. glabrescent/
4. covered with papillae/

FRUIT: a ripened ovary and any other structures that are attached and ripen with it.

GLABROUS: without hairs.

GLABRESCENT initially hairy but becoming glabrous.

PAPILLAE: pimple like protuberances.

#441. Fruit surface <venation whether visible>/
1. reticulate/
2. appearing veinless/
3. ribbed/

RETICULATE: net veined.

RIBBED: prominent veins causing ridges usually lengthwise.

#442. Capsule mouth <whether curved>/
1. straight/
2. curved/

#443. Capsule teeth <number of>/

#444. Legume <number of locules>/
1. unilocular/
2. nearly 2-locular by intrusion of placenta/

#445. Legume valves <whether twisted>/
1. twisted/
2. straight /

#446. Loment margins <whether winged>/
1. wingless/
2. winged with auricles/

#447. Styles <whether modifying and persisting>/
1. modified and persisting/
2. may persist until dehiscence but are not modified /

In some plants, the style changes structure in the fruiting stage to accommodate the dispersal of seeds, as in mountain avens. In other plants the style whithers and may prsist but does not assist seed dispersal.

STYLE: the narrowed portion of the pistil (gynoecium) connecting the stigma to the ovary.

CAUDUCOUS: the style falling early before the fruit is mature, or present and withered.

#448. Styles <shape of persisting style in fruit>/
1. becoming hooked/
2. becoming plumose/
3. remaining straight/

STYLE: the narrowed portion of the pistil (gynoecium) connecting the stigma to the ovary.

HOOKED: curved or with barbs that assist with animal dispersal.

PLUMOSE: feathery, or fluffy.

#449. Styles persisting in fruit <length in fruit>/
mm long/

Achenes in Cyperaceae

#450. Achenes <size relative to perigynia>/
1. filling the perigynia/
2. not filling the upper part of the perigynia/

ACHENE a small, nut-like fruit.

#451. Achenes <general shape>/
1. lenticular/
2. trigonous/
3. oblong/
4. subterete/
5. ovoid/
6. ovate/

LENTICULAR: Lens-shaped.

TRIGONOUS: a body triangular in cross section but with obtuse angles.

OBLONG: in the shape of a rectangle

SUBTERETE: approaching circular in cross section.

OVOID: egg-shaped.

OVATE: egg-shaped in outline and attached at the broad end.

Seeds

#452. Seeds <number per gynoecium>/

SEED: a ripened ovule. A seed has a single scar where it was attached to the placenta.

#453. Seeds <length>/
mm long/

SEED: a ripened ovule. A seed has a single scar where it was attached to the placenta.

#454. Seeds <colour>/
1. black/
2. brown/
3. white/
4. yellowish/

SEED: a ripened ovule.

#455. Seeds <surface>/
1. smooth/
2. hairy/
3. spinose/
4. verrucose/
5. ridged/
6. winged/
7. reticulate/
8. tuberculate/

SEED: a ripened ovule.

SMOOTH: an even, glabrous surface.

SPINOSE: with spines, which are stiff, slender, sharp-pointed structures arising from below the epidermis.

VERRUCOSE: covered with small wart-like rounded processes that are wider than high.

RIDGED: with folds (ridges).

WINGED: possessing wing-like structures.

RETICULATE: with a network pattern.

TUBERCULATE: covered with small wart-like rounded processes that are higher than wide.

Cytology

#456. 2n =/

#457. <Chromosome number reference: author and date>/

#458. Supposed basic chromosome number of family/

#459. Ploidy levels recorded/

Distribution

#460. <Northern hemisphere distribution>/
1. circumpolar/
2. circumboreal/
3. amphi-Atlantic/
4. amphi-Beringian/
5. North American/
6. not circumpolar, amphi-Atlantic, amphi-Beringian, or North American /

CIRCUMPOLAR: occurring in the area near and surrounding one of the poles of the earth.

AMPHI-ATLANTIC: occurring on both sides of the Atlantic Ocean.

AMPHI-BERINGIAN: occurring on both sides of the Bering Sea.

NORTH AMERICAN: occurring in the Arctic region in North America, Greenland, or Iceland.

#461. <North American phytogeography>/
1. arctic <widespread>/
2. high arctic <mainly restricted to the islands>/
3. low arctic <mainly restricted to the continent>/
4. alpine/
5. boreal/
6. coastal <mainly restricted to the coasts>/

#462. Range in the Canadian Arctic Archipelago/
1. widespread/
2. limited/
3. moderate /

#463. <Abundance in Arctic Islands>/
1. common/
2. uncommon/
3. rare/

#464. <Northern hemisphere: distribution>/
1. Greenland/
2. St. Pierre and Miquelon/
3. Canada/
4. United States/
5. Eurasia/

#465. Canada: <distribution>/
1. Alta./
2. B.C./
3. Lab./
4. Man./
5. N.B./
6. Nfld./
7. N.W.T. (Frank.)/
8. N.W.T. (Keew.)/
9. N.W.T. (Mack.)/
10. N.S./
11. Ont./
12. P.E.I./
13. Que./
14. Sask./
15. Yukon/
16. N.W.T. <after Nunavut was separated>/
17. Nunavut/

#466. <Distribution in the Canadian Arctic>/
1. Alaska/
2. Yukon/
3. North-west Territory Islands/
4. Continental North-west territories/
5. Nunavut Islands/
6. Continental Nunavut/
7. Northern Quebec/

#467. USA: <distribution>/
1. Alaska/
2. Ariz./
3. Calif./
4. Colo./
5. Idaho./
6. Maine/
7. Mich./
8. Minn./
9. Mont./
10. Nev./
11. N.H./
12. N.Mex./
13. N.York/
14. Oreg./
15. S. Dak./
16. Utah/
17. Vt./
18. Wash./
19. Wyo./

#468. Arctic Islands: <distribution>/
1. Baffin/
2. Devon/
3. Ellesmere/
4. Axel Heiberg/
5. Parry Islands <Melville, Bathurst, and Prince Patrick>/
6. Conrwallis/
7. Banks/
8. Victoria/
9. Prince of Wales/
10. Somerset/
11. King William/
12. Southampton/
13. Coats/
14. <small islands> /

#469. <Description of distribution>/

Ecology and Habitat

#470. Elevation/
m/

#471. Substrates, <habitat in the Arctic Islands>/
1. wet meadows/
2. hummocks <frost boils>/
3. snow beds/
4. around the margins of ponds/
5. depressions of low centre polygons/
6. marshes/
7. along streams/
8. river terraces/
9. lake shores /
10. tundra/
11. slopes/
12. ridges/
13. cliffs/
14. seashore/
15. dry meadows/

SNOWPATCH: The snowpatch is a more arctic plant habitat than the snowbed. It is usually a rather shallow depression in the landscape, where owing to the prevailing wind a snowdrift forms regularly each winter, affording protection for the plant cover beneath it. The snowdrift melts early and does not keep the habitat moist throughout the growing season. In the high-arctic landscape, plants with woody aerial stems such as willows, ground birch and heath are found chiefly on snowpatch habits. A snowbed is more usual in an arctic-alpine situation, where owing to topographical features, large masses of snow accumulate each winter. The plants growing on snowbed habitats enjoy the protection afforded by the snow cover and are assured of continuing water supply from the melting snow throughout the growing season but must be adapted to an even shorter growing season than those occupying more exposed habitats. In unfavourable seasons the snowbed habitat may remain covered by snow so late that the plants growing there may not have sufficient time to ripen their fruits, or they may not even have time to flower. For this reason only certain species of plants, can successfully occupy snowbed habitats.

#472. Substrates, <drainage>/
1. aquatic/
2. imperfectly drained moist areas/
3. on seepage slopes/
4. on solifluction slopes/
5. dry/
6. moderately well drained/

AQUATIC: growing in water.

#473. Substrates, <ionic character>/
1. acidic/
2. calcareous/
3. halophytic/
4. nitrophilous/
5. non-calcareous rocks/
6. non-littoral/

ACIDIC: growing in acidic soils.

CALCAREOUS: limestone derived alkaline soil.

HALOPHYTICY: soil with high salt content.

NITROGENOUS: soil enriched with nitrogen; e.g. old dung pile.

NON-LITTORAL: preferring drier conditions, usually away from the coast.

#474. Substrates, <particle size>/
1. rock/
2. gravel/
3. sand/
4. silt/
5. clay/
6. till/
7. moss/

#475. Substrates, <organic content>/
1. with low organic content/
2. with high organic content/
3. peat/

#476. <Notes:> /

Taxon as an Environmental Indicator

#477. <Taxon as an environmental indicator:>/

Indigenous Knowledge

#478. <Indigenous knowledge:>/

Economic Uses

#479. <Economic uses:>/

Notes

#480. <General notes:>/

#481. <Brief notes (extracted from ‘general notes’):>/

Metadata = miscellaneous

#482. <Number of> species in world/

#483. <Number of> genera in world/

#484. <Names of> genera in study region:/

#485. Morphological data <whether coded>/
1. coded /
2. not coded/
3. coding in progress/

#486. <Taxonomic level>/
1. species /
2. genus/
3. family/
4. class/

#487. <Whether accepted in this study>/
1. accepted in this study /
2. not accepted in this study <or, a synonym>/
3. status under consideration <or, not yet recorded on Arctic Islands, probably in the PAF Arctic area>/

Illustrations

#488. <Illustrations>/

Code used to indicate subject of image:

A = anatomy

B = black and white drawing

E = environment or habitat

C = close up of plant

F = flower

I = inflorescence

L = leaf or leaves

M = map

P = plate, where more than one picture is combined into a plate

R = roots

S = herbarium specimen

T = type specimen

U = fruit

X = cimage graphic

Working Comments

#489. <Working comments>/

#490. <Date, source material>/

This character gives the date when the data were gathered and the source of the data. Data for some species were based on published descriptions (citations given) supplemented by herbarium specimens. Others are based completely on original observations. The herbaria and sometimes states or provinces mainly sampled are given.

#491. Section used by Porsild (1957):/

#492. Code used for graphics files:/

#493. <Whether to be released at this time>/
1. to be released /
2. not to be released/

#494. PAF: <comments on name, September 2000>/

Classification

#495. Subclass/
1. Dicotyledonae <Magnoliopsida>/
2. Monocotyledonae <Liliopsida>/

#496. Dahlgren’s Superorder <of Dicotyledonae-modified from Dahlgren 1980)>/
1. Magnoliiflorae/
2. Ranunculiflorae/
3. Caryophylliflorae/
4. Polygoniflorae/
5. Malviflorae/
6. Primuliflorae/
7. Rosiflorae/
8. Fabiflorae/
9. Myrtiflorae/
10. Rutiflorae/
11. Asteriflorae/
12. Solaniflorae/
13. Corniflorae/
14. Gentianiflorae/
15. Lamiflorae/
16. Nymphaeiflorae/
17. Violflorae/

#497. Dahlgren’s Superorder <of Monocots: Dahlgren, Clifford and Yeo 1985>/
1. Alismatiflorae/
2. Liliiflorae/
3. Commeliniflorae/
4. Bromeliiflorae/

#498. Dahlgren’s Order <of Ranunculiflorae>/
1. Ranunculales/
2. Papaverales/

#499. Dahlgren’s Order <of Caryophylliflorae>/
1. Caryophyllales/

#500. Dahlgren’s Order <of Malviflorae>/
1. Plumbaginales/

#501. Dahlgren’s Order <of Violiflorae>/
1. Salicales/
2. Capparales/

#502. Dahlgren’s Order <of Primuliflorae>/
1. Primulales/

#503. Dahlgren’s Order <of Rosiflorae>/
1. Fagales/
2. Juglandales/
3. Saxifragales/
4. Rosales/

#504. Dahlgren’s Order <of Fabiflorae>/
1. Fabales/

#505. Dahlgren’s Order <of Myrtiflorae>/
1. Myrtales/
2. Haloragales/

#506. Dahlgren’s Order <of Rutiflorae>/
1. Geraniales/

#507. Dahlgren’s Order <of Solaniflorae>/
1. Solanales/
2. Boraginales/

#508. Dahlgren’s Order <of Asteriflorae>/
1. Campanulales/
2. Asterales/

#509. Dahlgren’s Order <of Corniflorae>/
1. Ericales/

#510. Dahlgren’s Order <of Gentianiflorae>/
1. Gentianales/

#511. Dahlgren’s Order <of Lamiiflorae>/
1. Scrophulariales/
2. Hippuridales/

#512. Dahlgren’s Order <of Alismatiflorae: Dahlgren, Clifford and Yeo 1985>/
1. Zosterales/

#513. Dahlgren’s Order <of Liliiflorae: Dahlgren, Clifford and Yeo 1985>/
1. Liliales/
2. Orchidales/

#514. Dahlgren’s Order <of Commeliniflorae: Dahlgren, Clifford and Yeo 1985>/
1. Cyperales/
2. Poales/

#515. Cronquist’s Subclass <of Magnoliopsida: 1981>/
1. Magnoliidae/
2. Hamamelidae/
3. Caryophyllidae/
4. Dilleniidae/
5. Rosidae/
6. Asteridae/

#516. Cronquist’s Order <of Magnoliidae>/
1. Ranunculales/
2. Papaverales/

#517. Cronquist’s Order <of Hamamelidae>/
1. Fagales/

#518. Cronquist’s Order <of Caryophyllidae>/
1. Caryophyllales/
2. Polygonales/
3. Plumbaginales/

#519. Cronquist’s Order <of Dilleniidae>/
1. Dilleniales/
2. Salicales/
3. Capparales/
4. Ericales/
5. Diapensiales/
6. Primulales/

#520. Cronquist’s Order <of Rosidae>/
1. Rosales/
2. Fabales/
3. Haloragales/
4. Myrtales/
5. Linales/

#521. Cronquist’s Order <of Asteridae>/
1. Gentianales/
2. Solanales/
3. Lamiales/
4. Callitrichales/
5. Plantaginales/
6. Scrophulariales/
7. Campanulales/
8. Asterales/

#522. Takhtajan’s Subclass <of Magnoliopsida: 1980>/
1. Ranunculidae/
2. Hamamelidae/
3. Caryophyllidae/
4. Dilleniidae/
5. Rosidae/
6. Asteridae/

#523. Takhtajan’s Superorder <of Dicots>/
1. Magnolianae/
2. Rafflesianae/
3. Ranunculanae/
4. Hamamelidanae/
5. Caryophyllanae/
6. Plumbaginanae/
7. Dillenianae/
8. Ericanae/
9. Rosanae/
10. Myrtanae/
11. Rutanae/
12. Asteranae/
13. Gentianaceae/
14. Laminae/

#524. Takhtajan’s Order <of Ranunculidae>/
1. Ranunculales/
2. Papaverales/

#525. Takhtajan’s Order <of Hamamelididae>/
1. Trochodendrales/
2. Fagales/
3. Cercidiphyllales/
4. Eupteleales/
5. Didymelales/
6. Hamamelidales/
7. Eucommiales/
8. Urticales/
9. Barbeyales/
10. Casuarinales/
11. Balanopales/
12. Leitneriales/
13. Myricales/
14. Juglandales/

#526. Takhtajan’s Order <of Caryophyllidae>/
1. Caryophyllales/
2. Polygonales/
3. Plumbaginales/

#527. Takhtajan’s Order <of Dilleniidae>/
1. Dilleniales/
2. Capparales/
3. Salicales/
4. Ericales/
5. Primulales/

#528. Takhtajan’s Order <of Rosidae>/
1. Saxifragales/
2. Rosales/
3. Fabales/
4. Myrtales/
5. Geraniales/

#529. Takhtajan’s Order <of Asteridae>/
1. Gentianales/
2. Polemoniales/
3. Scrophulariales/
4. Campanulales/
5. Asterales/


Cite this publication as: S.G. Aiken, M.C. LeBlanc, and M.J. Dallwitz 2000 onwards. Pteridophytes of the Canadian Arctic Archipelago: Descriptions, Illustrations, Identification, and Information Retrieval.’ Version: 23rd February 2001. http://www.mun.ca/biology/delta/arcticf/’. Dallwitz (1980), Dallwitz, Paine and Zurcher (1993, 1995, 2000) , and Aiken, Dallwitz et al. (1999) should also be cited (see References).

Index